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| Jepson eFlora
Key to families | Table of families and genera This text currently parallels The Jepson Manual: Vascular Plants of California, Second Edition that is now available at the University of California Press. Text appearing in blue on this page will not appear in the printed book; it will be displayed only on the Web. Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link. |
Annual to perennial herb ( subshrub).
Stem: generally erect.
Leaf: alternate, generally sessile, generally entire (toothed or lobed to ternately dissected).
Inflorescence: heads generally radiate (discoid, disciform), 1–few (many), peduncled; inflorescence generally ± flat-topped ( raceme- to panicle-like); involucre urn- to bell-shaped or generally hemispheric; phyllaries linear to narrowly lanceolate, in 2–several series, ± equal to strongly graduated, generally ascending or erect in flower, generally green, spreading when pressed, reflexed when dry; receptacle flat to steeply conic, smooth to shallowly pitted, epaleate.
Ray or pistillate flower: (0)10–generally many; ray generally narrow, generally white or pink to lavender or blue-purple (yellow), generally spreading when fresh, often coiled or reflexed when dry.
Disk flower: generally many; corolla generally narrowly funnel-shaped, yellow; anther tip ± lanceolate; style tips 0.1–0.8 mm, ± triangular.
Fruit: generally 0.5–3 mm, generally ± oblong, compressed to ± cylindric, generally 2-ribbed, generally sparsely hairy; pappus (0) generally double, outer of short bristles, narrow scales, or a short crown, inner of 6–50 long bristles.
± 375 species: worldwide. (Greek: early old age) [Nesom 2006 FNANM 20:256–348] Erigeron concinnus (Hook. & Arn.) Torr. & A. Gray var. condensatus D.C. Eaton, Erigeron disparipilus Cronquist, and Erigeron lobata A. Nelson apparently not in CA.
Unabridged references: [Noyes 2000 Plant Syst Evol 220:93–114]
1. Heads discoid, disciform, or inconspicuously radiate, rays 0 to barely exceeding involucre
2. Pistillate flowers present
3. Pistillate flowers of 2 kinds, those in outer zone with rays 3–4.5 mm, those in inner zone without rays ..... E. nivalis (2)
3' Pistillate flowers ± all alike, generally not in 2 zones
4. Heads 1; stem simple
5. Annual or biennial; pappus bristles 6–9(12) ..... E. divergens (3)
5' Perennial; pappus bristles 12–25
6. Lvs (1)2–3(4)-ternately dissected ..... E. compositus (2)
6' Lvs entire
7. St minutely glandular; disk corolla generally abruptly inflated to throat; fruit base ringed ± white; outer pappus of narrow scales or obvious bristles ..... E. aphanactis var. congestus
7' St glandless; disk corolla narrowly funnel-shaped; fruit base ringed ± yellow; outer pappus of minute bristles ..... E. chrysopsidis var. austiniae
4' Heads 1–many; stem often branched
8. Rays 1–3 mm
9. Infl panicle-like, heads many; rays <= 1 mm; involucre glabrous or sparsely strigose to minutely bristly; central colored part of middle phyllaries resin-filled, ± red-brown when dry ..... E. canadensis (3)
9' Infl generally raceme-like, heads 1–12; rays 2–3 mm; involucre ± rough-hairy; central colored part of middle phyllaries not resin-filled, not red-brown when dry ..... E. lonchophyllus (2)
8' Rays 0–1 mm
10. Central stem and axes of stiffly ascending branches generally strongly dominant; inflorescence generally raceme- or panicle-like, often many-headed; disk flowers 6–25
11. Central stem often overtopped by distal branches in age; plant ± gray-hairy; heads ± long-peduncled, generally in raceme-like clusters; receptacle (2.5)3–5 mm diam in fruit; pistillate flowers 60–150+; phyllaries generally minutely bristly and/or densely strigose; pappus 3–4+ mm ..... E. bonariensis
11' Central stem generally exceeding distal branches in age; plant green, glabrous to hairy; heads ± short-peduncled in panicle-like clusters; receptacle 1–3 mm diam in fruit; pistillate flowers 20–90+; phyllaries glabrous to densely strigose or densely bristly; pappus 2–3(4) mm
12. Lf ± long-bristly-ciliate; phyllaries generally glabrous, occasionally sparsely strigose; central colored part of middle phyllaries narrower than light-colored margin, resin-filled, ± red-brown when dry; ray 0.3–1 mm; disk corolla generally 4-lobed; fruit uniformly pale tan to light gray-brown ..... E. canadensis (3)
12' Lf margin generally ± fine-strigose; phyllaries glabrous to densely strigose or spreading-hairy; central part of middle phyllaries generally wider than lighter margin, not red-brown or resin-filled when dry; ray 0–0.3 mm; disk corolla generally 5-lobed; fruit pale tan, generally some in head with ± red veins ..... E. sumatrensis
10' Central stem and axes of ascending branches not strongly dominant, plant ± openly branched; inflorescence ± cyme-like, often 1–few-headed; disk flowers generally >= 25
13. Perennial; pappus bristles 12–17 ..... E. aphanactis var. aphanactis
13' Annual, biennial or short-lived perennial herb
14. Pappus bristles 15–20; biennial or short lived per ..... E. calvus
14' Pappus bristles 6–9(12); annual or biennial ..... E. divergens (3)
2' Pistillate flowers 0
15. Basal leaves present at flower; cauline leaves generally much-reduced or 0 distally
16. Caudex simple ..... E. lassenianus var. deficiens
16' Caudex branched
17. St ascending, 15–40 cm, occasionally sparsely spreading-hairy; caudex branches slender, rhizome-like; basal leaves elliptic to oblanceolate to spoon-shaped ..... E. supplex
17' St erect, 4–15 cm, ± glabrous to densely strigose; caudex branches short, stout; basal leaves ± linear ..... E. bloomeri
18. Herbage and phyllaries strigose ..... var. bloomeri
18' Herbage and phyllaries ± glabrous ..... var. nudatus
15' Basal leaves 0 at flower; cauline leaves generally evenly sized and spaced
19. Glandless stem hairs ± spreading
20. St hairs dense, short, stiff, slightly reflexed, ± gray ..... E. inornatus var. keilii
20' St hairs sparse to dense, long, ± crinkly-spreading, white
21. St 30–90 cm, ascending to erect; leaves 20–40 mm; involucre 12–15 mm diam ..... E. biolettii (2)
21' St 5–30 cm, decumbent to ascending; leaves 7–25 mm; involucre 7–15 mm diam
22. Inner phyllaries 3.5–5 mm ..... E. miser
22' Inner phyllaries 5.5–7 mm ..... E. petrophilus
23. Herbage ± glandless; hairs stiff, straight or curved ..... var. viscidulus
23' Herbage densely glandular; glandless hairs generally loose, often crinkly
24. Phyllary tips obviously widened and ± purple ..... var. petrophilus
24' Phyllary tips not widened or purple ..... var. sierrensis
19' Glandless stem hairs 0–sparse, appressed
25. Phyllaries generally glabrous ..... E. inornatus (in part)
26. St 10–20 cm, decumbent to ascending; proximally ± reflexed-hairy; proximal and middle leaf blades and margins stiffly spreading-hairy and -ciliate; inner phyllaries 5–6.5 mm ..... var. calidipetris
26' St 30–90 cm, ascending; proximally ± glabrous or appressed-hairy; proximal and middle leaf blades and margins generally short-strigose or ± glabrous; inner phyllaries 4.5–5.5 mm ..... var. inornatus
25' Phyllaries glandular (sometimes also with sparse, glandless hairs)
27. Herbage glandular; leaf narrowly oblanceolate ..... E. biolettii (2)
27' Herbage glandless; leaf thread-like to oblanceolate
28. Lf ± oblanceolate; stem long-ascending from woody taproot ..... E. inornatus var. inornatus
28' Lf thread-like to linear; stem from stout woody caudex or thin, rhizome-like caudex branches
29. St 30–90 cm, many, from stout woody caudex ..... E. greenei
29' St 8–20(30) cm, 1, from slender, rhizome-like caudex branches ..... E. reductus
30. Phyllary tips generally ± purple (variable n SnFrB); pappus bristles 38–61 ..... var. angustatus
30' Phyllary tips ± green; pappus bristles 20–30 ..... var. reductus
1' Heads conspicuously radiate, rays exceeding to much-exceeding involucre
31. Pistillate flowers of 2 kinds, those in outer zone with rays 3–4.5 mm, those in inner zone without rays ..... E. nivalis (2)
31' Pistillate flowers all alike, not in 2 zones
32. Rays 1–3 mm
33. Infl panicle-like, heads many; rays <= 1 mm; involucre glabrous or sparsely strigose to minutely bristly; central colored part of middle phyllaries resin-filled, ± red-brown when dry ..... E. canadensis (3)
33' Infl generally raceme-like, heads 1–12; rays 2–3 mm; involucre ± rough-hairy; central colored part of middle phyllaries not resin-filled, not red-brown when dry ..... E. lonchophyllus (2)
32' Rays 4–25 mm
34. Annual (biennial to short-lived perennial herb) from slender taproot or shallow fibrous roots (see also Erigeron philadelphicus var. philadelphus)
35. Ray flower pappus 0; plant fibrous-rooted
36. Lvs generally coarsely toothed; stem hairs sparsely spreading or appressed ..... E. annuus
36' Lvs generally entire; stem hairs appressed ..... E. strigosus var. strigosus
35' Ray flower pappus present; plant taprooted (Erigeron flagellaris generally fibrous-rooted)
37. St hairs appressed; plant with herbaceous, leafy runners ..... E. flagellaris
37' St hairs spreading; plant without runners ..... E. divergens (3)
34' Perennial or subshrub from woody taproot, branched caudex, or rhizome
38. Lvs 1–4-ternately dissected
39. Lvs generally (1)2–3(4)-ternately dissected; ray < 1 mm wide; caudex branches short, thick, ascending ..... E. compositus (2)
39' Lvs deeply 3-lobed at tip; ray 1–2 mm wide; caudex branches thin, rhizome-like ..... E. vagus
38' Lvs entire to shallowly toothed
40. Lf bases hardened, white-shiny, expanded, enclosing proximal stem
41. Caudex branches thick, leaf bases persistent; rays yellow ..... E. linearis
41' Caudex branches slender, rhizome-like, leaf bases not persistent; rays white, pink to blue or lavender
42. Involucre 13–18 mm diam; basal leaves generally 2–5 mm wide; pappus bristles 25–40; phyllary hairs spreading; >= 2100 m; CaRH, n&c SNH ..... E. barbellulatus
42' Involucre 6–11 mm diam; basal leaves generally 0.5–1 mm wide; pappus bristles 20–30; phyllary hairs appressed; 1000–2200 m; MP ..... E. elegantulus
40' Lf bases ± thin, not hardened, barely or not expanded
43. Fr ribs 4–8; herbage generally ± silvery, hairs dense, appressed
44. Fr ribs 6–8; basal leaves tufted, persistent ..... E. argentatus
44' Fr ribs 4(6); basal leaves generally 0 at flower
45. Outer pappus conspicuous, of bristles or narrow scales; stem silvery-white ..... E. parishii
45' Outer pappus inconspicuous; stem gray-green, densely strigose ..... E. utahensis
43' Fr ribs generally 2 (if more, herbage not silvery)
46. Phyllaries strongly graduated; leaves entire, not clasping, basal 0, cauline generally evenly sized and spaced
47. St minutely glandular ..... E. aequifolius
47' St generally not glandular
48. St hairs dense, minutely curled- or bent; fruit glabrous ..... E. blochmaniae
48' St hairs 0 to dense, straight, stiffly spreading to ± reflexed or appressed; fruit sparsely strigose
49. St hairs dense, stiffly spreading, generally slightly reflexed
50. Herbage hairs 0.5–1 mm; phyllaries densely minutely glandular, inner phyllary tips widely white-thick-margined; stem often from woody basal offsets and roots ..... E. klamathensis
50' Herbage hairs <= 0.1–0.4 mm; phyllaries glandular or glandless, inner phyllary tips green; stem from ± slender, rhizome-like bases ..... E. breweri
51. Phyllaries densely glandular, glandless hairs 0 or very sparse
52. St generally >= 20 cm, ascending to erect; leaves generally >= 15 mm ..... var. breweri
52' St 7–15 cm, prostrate to decumbent; leaves 5–12 mm ..... var. jacinteus
51' Phyllaries not or barely glandular, glandless hairs prominent
53. St 20–30 cm, wiry, brittle; phyllary hairs long, thick-based, stiffly spreading, translucent (also some glandular) ..... var. porphyreticus
53' St 30–75 cm, not wiry or brittle; phyllary hairs short, white, ± appressed, glandless
54. Phyllary hairs thin-based, ± appressed, slightly less dense on inner phyllaries ..... var. bisanctus
54' Phyllary hairs thick-based, ascending, much less dense on middle and inner phyllaries ..... var. covillei
49' St hairs ± 0 or sparse, appressed
55. St prostrate to ± erect, from slender rhizome ..... E. elmeri
55' St decumbent to erect, from woody taproot
56. Ray flowers 9–13; leaf ciliate, hairs thin-based; phyllaries with minute glandless hairs and densely minutely glandular ..... E. serpentinus
56' Ray flowers 15–60; leaf ciliate, hairs thick-based; phyllaries glandular or glandless
57. Phyllaries ± glandless, 0.5–0.8 mm wide, margins generally thick to ± scarious
58. Lvs 2–4(5) cm, oriented variously; inner phyllaries 3.2–4.5 mm ..... E. foliosus var. foliosus
58' Lvs (2.5)4–6(8) cm, often ± oriented to 1 side of stem; inner phyllaries (4)5–6 mm
59. St 30–85 cm; axillary leaf tufts 0; leaves 35–80 mm, 1–2(4) mm wide; inner phyllaries not strongly scarious; ray flowers 26–50 ..... E. foliosus var. hartwegii
59' St 15–28 cm; axillary leaf tufts generally present; leaves 25–45 mm, (2)5–8 mm wide; inner phyllary margins strongly scarious; ray flowers 18–22 ..... E. mariposanus
57' Phyllaries minutely but prominently glandular, 0.8–1 mm wide, margins generally widely scarious ..... E. foliosus (in part)
60. Phyllary hairs all glandular; leaves generally 1–2 mm wide ..... var. confinis
60' Phyllary hairs glandular and glandless, ± densely bristly-strigose; leaves generally 2–4 mm wide
61. Lf blades ± strigose; phyllary midvein raised, orange-resinous; ray corolla 7–10 mm ..... var. franciscensis
61' Lf blades ± glabrous (except ciliate hairs); phyllary midvein generally not raised or orange-resinous; ray corolla 10–15 mm ..... var. mendocinus
46' Phyllaries generally ± equal; leaves entire to lobed, sometimes clasping, basal sometimes present, cauline often strongly reduced distally on stem, if present
62. Plant fibrous-rooted, generally from obvious rhizome or ± decumbent caudex; cauline leaves often ± clasping;
63. Glandless phyllary hairs with black cross-walls (sometimes only on basal cells of hairs)
64. Rays white ..... E. coulteri
64' Rays blue to purple ..... E. sanctarum (2)
63' Glandless phyllary hairs 0 or with clear cross-walls
65. Ray flowers 150–400 ..... E. philadelphicus var. philadelphicus
65' Ray flowers 20–165
66. Lvs thick, ± fleshy, entire or toothed; coastal ..... E. glaucus
66' Lvs ± thin, entire; coastal or inland
67. Glandless phyllary hairs prominent, glandular hairs sometimes present
68. St from thin, often stolon-like caudex branches with few fibrous roots ..... E. sanctarum (2)
68' St from short, thick, strongly fibrous-rooted rhizome or caudex
69. St 2–20(30) cm, hairs spreading; phyllaries ± black-purple; pappus bristles 12–16(20) ..... E. algidus (2)
69' St 30–100 cm, hairs appressed; phyllaries ± green; pappus bristles 20–30 ..... E. aliceae (2)
67' Glandless phyllary hairs 0, glandular hairs dense
70. St glabrous or sparsely minutely glandular; ray flowers 20–30(45) ..... E. cervinus
70' St ± appressed-hairy, sometimes also ± bristly; ray flowers 30–105
71. Fr 2(4)-ribbed, moderately to densely strigose; proximal leaves generally coarsely dentate, sometimes entire ..... E. aliceae (2)
71' Fr 4–7-ribbed, sparsely strigose; proximal leaves entire ..... E. glacialis
72. Peduncles densely minutely strigose with loosely appressed, slightly crinkled hairs ..... var. glacialis
72' Peduncles bristly or ± stiffly long-soft-hairy ..... var. hirsutus
62' Plant taprooted or with taprooted caudex; cauline leaves not at all clasping
73. St spreading- or reflexed-hairy
74. Lf clearly petioled, blade oblanceolate, elliptic, obovate to spoon-shaped
75. Basal leaves 20–70 mm; cauline leaves much reduced; stem 2–30 cm; phyllaries ± black-purple; ray flowers (30)50–125, ray 7–13 mm ..... E. algidus (2)
75' Basal leaves 5–25 mm; cauline leaves 0; stem 1–4 cm; phyllaries green; ray flowers 15–40, ray 4–6 mm ..... E. uncialis var. uncialis
74' Petiole indistinct, blade linear or oblanceolate, gradually tapered to base
76. Margins of proximal leaves generally hairy but not stiffly spreading-ciliate; generally > 2200 m
77. Basal leaves 20–80 mm; stem sparsely minutely glandular at least distally, and moderately hairy with short, stiffly spreading to reflexed, glandless hairs; phyllaries green; ray flowers 25–55, ray 6–11 mm ..... E. clokeyi var. pinzliae
77' Basal leaves 6–35 mm; stem densely minutely glandular, also with short-stiff and ± spreading glandless hairs; phyllaries ± black-purple; ray flowers 20–37, ray 4–7(10) mm ..... E. pygmaeus
76' Margins of proximal leaves generally stiffly long-spreading-ciliate; 1200–1800 m
78. Disk corolla (±) glabrous; pappus bristles 12–22; outer pappus of inconspicuous bristles or narrow scales; rays (40)55–115, 8–12 mm ..... E. pumilus var. intermedius
78' Disk corolla sharply scabrous; pappus bristles 5–15; outer pappus of prominent wide scales; rays 40–60, 7–9 mm ..... E. concinnus var. concinnus
73' St glabrous or ascending- to appressed-hairy (to ± spreading-hairy in Erigeron lassenianus)
79. Ray flowers 75–125; disk corolla throat ± widened ..... E. multiceps
79' Ray flowers generally 14–80; disk corolla ± narrowly cylindric
80. Basal leaves 0 at flower
81. Lvs sharply reduced distally on stem; axillary leaf tufts 0; ray flowers 12–27(45) ..... E. oxyphyllus
81' Only most distal cauline leaves reduced; leaf tufts in axils; ray flowers 45–80 ..... E. karvinskianus
80' Basal leaves generally persistent
82. Basal leaves thread-like to linear
83. St 2–8 cm, leafless; stem hairs straight, appressed; basal leaves 5–25 mm; fruit ribs densely ciliate, faces glabrous ..... E. compactus
83' St 15–30 cm, generally leafy, at least proximally; stem hairs curved up, ± loose; basal leaves generally 20–70 mm; fruit ribs and faces sparsely hairy ..... E. filifolius
82' Basal leaves narrowly oblanceolate to elliptic or spoon-shaped (or linear in Erigeron lassenianus var. lassenianus and Erigeron robustior)
84. Plants cespitose; caudex stout, branched; basal leaves oblanceolate to elliptic, petioled; cauline leaves abruptly reduced ..... E. tener
84' Plants not cespitose, caudex generally slender; basal leaves linear to narrowly oblanceolate, petiole indistinct; cauline leaves gradually reduced
85. Basal leaves generally persistent; cauline leaves generally smaller than basal; heads 1–4(7) ..... E. eatonii
86. Involucre (14)17–23 mm diam; longest phyllaries 7–11 mm; disk corolla 4.4–6.8 mm ..... var. nevadincola
86' Involucre 8–16 mm diam; longest phyllaries 5–8 mm; disk corolla 3–5 mm
87. St 10–23 cm; disk corolla 3–4 mm; pappus bristles 16–20, 3–3.5 mm ..... var. plantagineus
87' St 4–12(21) cm; disk corolla 3.5–5 mm; pappus bristles 18–30, 3.5–5 mm ..... var. sonnei
85' Basal leaves persistent at, or withering before, flower; cauline leaves gradually or little reduced; heads (1)2–7(18)
88. Phyllaries densely minutely glandular; involucre 4–5.5 mm, 6–10(12) mm wide; leaves generally 1-veined ..... E. lassenianus var. lassenianus
88' Phyllaries generally glandless; involucre generally 6–8.5 mm, (9)12–20 mm wide; leaves generally 3-veined
89. Plants 10–22(27) cm; taproots ± thick, (3)5–8 mm wide; basal leaves oblanceolate to ± narrowly spoon-shaped; cauline leaves ± continuing to heads; involucres (5)6–7 mm, 9–12(14) mm wide; phyllaries elliptic-oblanceolate to oblong-oblanceolate, abruptly acuminate ..... E. maniopotamicus
89' Plants (15)25–55 cm; taproots ± thin, 2–3 mm wide; basal leaves linear to very narrowly oblanceolate; cauline leaves ending ± well below heads; involucres 6–8.5 mm, (12)14–20 mm wide; phyllaries narrowly oblanceolate to lanceolate, acute-acuminate ..... E. robustior
Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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