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Key to families | Table of families and genera

Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.



Robert L. Carr & Gerald D. Carr

Annual, erect to spreading, with coarse tack-shaped glands, and often simple glandular hairs, aromatic; generally self-sterile. Leaf: basal ± many, mostly deciduous by flowering, occasionally persistent, margins often curled under, and cauline, generally alternate and reduced distally on stem, ± sessile, ± linear, entire or sparsely toothed, ± scabrous, generally with longer stiff hairs, especially on proximal margins. Inflorescence: heads radiate, axillary, often ± sessile, and terminal, 1 or clustered; peduncle bracts linear or narrowly lanceolate to widely oblanceolate, margins thickened, rounded, defining a ± prominent central abaxial channel, especially proximally, ± bristly-ciliate, especially proximally, some or all with 1+ tack-like glands; phyllaries ± equal to shorter than paleae, each partly enfolding a ray fruit; receptacle ± flat, hairy; paleae in one series between ray and disk flowers, ± fused, forming a receptacle cup, phyllaries, paleae generally with abaxial simple and/or tack-like glands and appressed and/or spreading hairs. Ray flower: 1–6; ray 2–12 mm, 3-lobed, white to rose or yellow, occasionally with red spot near base. Disk flower: 1–25; corolla 2.5–10 mm, colored ± like rays; anthers generally purple, tips ovate to triangular; style branches long, bristly. Fruit: ray fruit 1.5–4 mm, ± 3-angled, adaxial face ± flat, abaxial face rounded to ± angled, pappus 0; disk fruit 2–4 mm, angled, base tapered, ± appressed-hairy to glabrous, pappus 0 or generally of 6–16 scales, all ± lanceolate or some blunt and shorter.
± 10 species: especially California (Oregon). (Greek: cup gland) [Carr & Carr 2006 FNANM 21:270–276]

Key to Calycadenia

1. Tack-like glands of peduncle bracts (0)1, terminal; central ray lobe generally << laterals, widest at base

2. Corolla yellow; ray fruit generally rough-wrinkled

3. Plant 1–5 dm, ± slender; ray 2–2.5(3.5) mm; disk flower 1–3 ..... C. micrantha

3' Plant 2–12 dm, occasionally robust; ray (4)5–12 mm; disk flower 3–25 ..... C. truncata

2' Corolla generally white, occasionally aging ± red; ray fruit generally smooth

4. Branchlets generally many, thread-like, very flexible, ray flower (0)1(2); disk flower 1–2; ray fruit ± glabrous ..... C. hooveri

4' Branchlets, if present, coarser, ± rigid, ray flower 1–5; disk flower 4–15; ray fruit ± densely appressed-hairy

5. Tack-like glands of phyllaries and paleae generally many, variable in size; disk flower corolla 7–10 mm ..... C. spicata

5' Tack-like glands generally 0 on phyllaries and paleae; disk flower corolla 5–6 mm ..... C. villosa

1' Tack-like glands of peduncle bracts generally 2–6+, terminal and/or scattered; central ray lobe <= laterals, generally widest near or beyond middle

6. Peduncle bracts ± widely oblanceolate, central channel ± not narrowed at tip, tip ± widely rounded, tack-like glands marginal, ± uniform in size; phyllaries and paleae generally without tack-like glands ..... C. mollis

6' Peduncle bracts ± linear or elliptic to lanceolate or club-like, central channel narrowed or 0 distally, tip acute, rounded, or truncate, tack-like glands variable in size and position; phyllaries and paleae often with tack-like glands

7. Central lobe of ray widest at base, << and narrower than laterals ..... C. multiglandulosa

7' Central lobe of ray widest near or beyond middle, ± similar to laterals

8. Leaves all opposite; heads in congested, axillary and terminal clusters, appearing whorled ..... C. oppositifolia

8' Leaves generally alternate; heads 1 or clustered in axils, in spike-like or diffusely branched clusters, not appearing whorled

9. Stem generally simple or few-branched, ± rigid, main axis generally obvious; paleae (3)4–6+, receptacle cup ± bell-shaped, length ± = diam; ray flower (1)2–6, ray yellow, cream, or white, aging ± pink; disk flower (4)6–21 ..... C. fremontii

9' Stem branched, main stem axis generally 0, branches ± many, slender, flexible, generally divergent, often zigzag; paleae 3(4), receptacle cup ± fusiform to club-shaped, length ± 2 × diam; ray flower 1(2), ray white, aging ± pink, never yellow; disk flower 2–5 ..... C. pauciflora


Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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