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Key to families | Table of families and genera

Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.

 

CALYSTEGIA MORNING-GLORY

R.K. Brummitt


Perennial herb, subshrub from caudex or rhizome, glabrous to tomentose. Stem: short to high-climbing, generally twisting, twining. Leaf: generally > 1 cm, linear to reniform or sagittate to hastate (deeply divided). Inflorescence: peduncle generally 1-flowered; bracts generally ± opposite, lobed or not, > 1 mm below calyx, not hiding it, small, to < 1 mm below calyx, hiding it or ± so, large. Flower: generally showy; corolla glabrous, white or yellow to pink or purple; ovary chamber 1, style 1, stigma lobes 2, oblong, tips obtuse. Fruit: ± spheric, ± inflated. Seed: generally ± 4.
± 25 species: temperate, worldwide. (Greek: hiding calyx, by bracts of some) [Brummitt 2002 Madroño 49:130–131] Intermediates common, often difficult to identify. Molecular evidence indicates close relationship with Convolvulus (Carine et al. 2004 Amer J Bot 91:1070–1085). Bracts qualify as bractlets by some definitions. Leaf blade length measured along midrib.
Unabridged references: [Brummitt 1980 Kew Bull 35(2):327–328]

Key to Calystegia

1. Bracts > 1 mm below calyx, not hiding it, <= 5 mm wide or variously toothed or lobed like leaves

2. Plant glabrous

3. Bracts elliptic to widely elliptic-oblong, entire ..... C. peirsonii (2)

3' Bracts generally linear, entire or lobes basal

4. Stem ± stiffly erect, intertwined; leaf linear to narrowly triangular, lobes 0 or ± linear, sinus rounded ..... C. longipes

4' Stem trailing to strongly climbing, ± not intertwined; leaf triangular to reniform, lobes wider than linear, 2(3)-tipped, sinus V-shaped or ± closed ..... C. purpurata

5. Stem strongly climbing, > 1 m; leaf triangular, tip acute, sinus V-shaped, bracts rarely lobed ..... subsp. purpurata

5' Stem trailing to weakly climbing, generally < 1 m; leaf ovate-triangular to reniform, tip generally rounded to notched, sinus generally ± closed; bracts generally with small lobes ..... subsp. saxicola

2' Plant puberulent to tomentose at least around leaf sinus or at peduncle tip

6. Bracts entire, not lobed like leaves, ± sessile ..... C. occidentalis subsp. occidentalis

6' Bracts toothed or lobed like leaves, stalked

7. Leaf lobes 7–9, linear, finger-like ..... C. stebbinsii

7' Leaf ± entire or lobes 2, not linear, not finger-like

8. Leaf blade widely triangular to ± reniform, tip acute to acuminate or notched, lobes generally 2-tipped; hairs ± densely spreading ..... C. malacophylla subsp. malacophylla

8' Leaf blade generally narrowly triangular, tip ± acute, lobes generally 1-tipped; hairs minute or fine ..... C. occidentalis subsp. fulcrata

1' Bracts < 1 mm below calyx, ± hiding it, often > 4 mm wide, not lobed like leaves

9. Bracts < 4 mm wide

10. Plant hairs dense, not appressed ..... C. collina subsp. tridactylosa

10' Plant hairs 0 or minutely appressed

11. Leaf blade narrowly triangular, lobes clearly defined, directed ± basally, base with deep sinus ..... C. peirsonii (2)

11' Leaf blade generally triangular-hastate to reniform, lobes not clearly defined, directed ± laterally, base truncate to wedge-shaped or widely rounded, occasionally with shallow sinus

12. Bracts ± obtuse; plant glabrous ..... C. atriplicifolia subsp. buttensis (2)

12' Bracts acute; plant hairs generally minutely appressed ..... C. subacaulis subsp. episcopalis (2)

9' Bracts > 4 mm wide

13. Stem trailing or weakly to strongly climbing, >= 1 m

14. Plant from rhizomes, stem not woody; peduncle 1-flowered; marshes, riverbanks, or disturbed areas

15. Bracts strongly sac-like at base, margins overlapped; corolla generally 55–75 mm ..... C. silvatica subsp. disjuncta

15' Bracts not strongly sac-like at base, margins not overlapped; corolla 30–58 mm ..... C. sepium

16. Bracts < sepals; corolla generally ± 40 mm ..... subsp. binghamiae

16' Bracts generally > sepals; corolla 30–58 mm ..... subsp. limnophila

14' Plant from woody caudex, lower stem often woody; peduncle often > 1-flowered; generally dry places ..... C. macrostegia

17. Bracts 16–30 mm wide, strongly keeled to ± sac-like at base

18. Bracts 19–37 mm; calyx 16–25 mm; corolla 47–68 mm; stamens 23–35 mm; s Channel Islands ..... subsp. amplissima

18' Bracts 13–26 mm; calyx 10–22 mm; corolla 36–60 mm; stamens 17–26 mm; n Channel Islands ..... subsp. macrostegia

17' Bracts 4–16 mm wide, flat or ± keeled

19. Bracts obtuse ..... subsp. cyclostegia

19' Bracts acute

20. Leaves hairy ..... subsp. arida

20' Leaves glabrous to puberulent

21. Leaf blade triangular, > 7 mm wide except basal lobes ..... subsp. intermedia

21' Leaf blade linear to narrowly triangular, < 7 mm wide except basal lobes ..... subsp. tenuifolia

13' Stem not or only weakly climbing, < 1 m

22. Plant hairs 0 or sparsely appressed-minute and leaf triangular-hastate to reniform

23. Corolla deep pink or ± purple; leaf reniform, glabrous, ± fleshy; ocean beaches, dunes ..... C. soldanella

23' Corolla white or ± yellow (pink- or purple-tinged); leaf ± reniform or not, glabrous or not, not fleshy; inland

24. Bracts obtuse; plant generally glabrous ..... C. atriplicifolia subsp. buttensis (2)

24' Bracts acute; plant generally minutely appressed-hairy ..... C. subacaulis subsp. episcopalis (2)

22' Plant tomentose or hairs densely spreading, or leaf not triangular-hastate to reniform

25. Stem well developed, < 100 cm, without basal rosette of leaves

26. Plant tomentose; leaf margin wavy ..... C. collina subsp. venusta (2)

26' Plant tomentose or hairs densely spreading; leaf margin not wavy ..... C. malacophylla

27. Leaf widely triangular to ± reniform, tip generally obtuse to notched, lobes generally 2-tipped; hairs brown or golden-brown (± gray); Cascade Range, Sierra Nevada ..... subsp. malacophylla

27' Leaf ± narrowly triangular, tip generally acute, lobes generally 1-tipped (2nd tip 0 or poorly developed); hairs ± gray (± brown); San Francisco Bay Area, South Coast Ranges, Western Transverse Ranges ..... subsp. pedicellata

25' Stem poorly developed, < 30 cm, <= basal rosette of leaves

28. Plant hairs sparse to ± dense, short, spreading to reflexed; leaf margin not or ± wavy ..... C. subacaulis subsp. subacaulis

28' Plant ± tomentose; leaf margin wavy ..... C. collina

29. Sepals tomentose ..... subsp. venusta (2)

29' Sepals glabrous or strigose at center

30. Leaf tip obtuse, lobes generally indistinct, blade triangular to reniform ..... subsp. collina

30' Leaf tip acute, lobes distinct, blade triangular ..... subsp. oxyphylla

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