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Key to families | Table of families and genera
This text currently parallels The Jepson Manual: Vascular Plants of California, Second Edition that is now available at the University of California Press.
Text appearing in blue on this page will not appear in the printed book; it will be displayed only on the Web. Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.
Shrub to vine, often thicket-forming, generally prickly.
Leaf: generally odd-pinnately compound; stipules generally attached to petiole, generally gland-margined.
Inflorescence: generally ± cyme or flowers 1; pedicel bractlets 0.
Flower: hypanthium urn-shaped, bractlets 0; sepals often with long expanded tip; petals generally 5 (except cultivated), generally pink in CA (white to red or yellow); stamens generally > 20; pistils generally many, ovaries superior, styles attached at tip, generally hairy.
Fruit: bony achenes generally enclosed in fleshy, generally ± red hypanthium (hip).
100+ species: generally n temperate. (Latin: ancient name) [Ertter & Lewis 2008 Madroño 55:170–177] Spp. hybridize freely; other non-natives established locally. FNANM treatment by Lewis & Ertter uses both subspp. and vars., the latter mostly reserved for localized variants within a subsp.
Unabridged references: [Lewis & Ertter 2007 Novon 17:342–353]
1. Lflets << 1 cm, toothed ± 1/2 to base; hypanthium densely prickly — PR ..... R. minutifolia
1' Lflets generally > 1 cm, toothed < 1/4 to base; hypanthium glabrous to stalked-glandular
2. Sepals with toothed lateral lobes; prickles compressed side-to-side, curved
3. Hypanthium 1–2 mm wide at flower, 5–7 mm wide at fruit; pistils generally < 10 ..... R. multiflora
3' Hypanthium 4–5 mm wide at flower, 10–20 mm wide at fruit; pistils > 10
4. Lvs and sepals ± glandless; petals white to pink; terminal leaflet ± ovate ..... R. canina
4' Lvs and sepals glandular; petals pink; terminal leaflet generally elliptic to ± widely obovate ..... R. rubiginosa
2' Sepals generally entire (tip sometimes toothed); prickles compressed or slender, straight or curved
5. Sepals deciduous in fruit; hypanthium 1.5–2 mm wide at flower; pistils 5–10; sepal tip generally << body; petals ± 10 mm — leaflets glabrous, flowers 1–3; pedicels generally stalked-glandular ..... R. gymnocarpa
6. Lflets generally (5)7–9; terminal leaflet tip generally ± obtuse, ± 10–30 mm; pedicels ± 15–30 mm; plants to ± 5–20 dm, generally in shade on non-ultramafic substrates; widespread ..... var. gymnocarpa
6' Lflets generally 5(7); terminal leaflet tip widely obtuse to rounded, 4–20 mm; pedicels ± 10–15 mm; plants to generally 3–6 dm, full sun on ultramafic substrates; KR ..... var. serpentina
5' Sepals persistent; hypanthium 2.5–7 mm wide at flower; pistils generally > 10; sepal tip ± = or often > body; petals 10–25 mm
7. Dwarf, openly rhizomed shrubs, generally 1–5(10) dm; leaflet tip often ± truncate — leaf margins generally ± glandular, generally double-toothed
8. Hypanthium stalked-glandular — prickles generally ± slender, often many; NW, CW, MP(1 site), 150–1550(1950) m ..... R. spithamea
8' Hypanthium generally glandless, rarely sparsely glandular
9. Prickles few, paired, ± thick-based; terminal leaflet generally widely obovate; flower 1–2(7); CaR, SN, 700–2500 m ..... R. bridgesii
9' Prickles generally many, generally not paired, both slender and ± thick-based; terminal leaflet generally ± elliptic; flowers generally 1–5; CW, generally < 300 m ..... R. pinetorum
7' Open or thicket-forming shrubs, generally > 5 dm; leaflet tip not truncate
10. Terminal leaflet ± obovate-elliptic, widest at or above middle; leaf axis finely velvety, hairs ± 0.1 mm (glabrous), glandless; prickles slender to ± thick-based, generally ± straight; sepals glandless (in CA); CaRH, SNH, Teh, TR, GB, DMtns ..... R. woodsii
11. Prickles generally many, 2–10(13) mm, often ± thick-based, internodal prickles generally present in inflorescence; c&s SNH, Teh, TR, SNE, DMtns ..... subsp. gratissima
11' Prickes few to many, 2–7 mm, slender, internodal prickles generally 0 in inflorescence; CaRH, n SNH, MP, n SNE ..... subsp. ultramontana
10' Terminal leaflet generally ovate to elliptic, generally widest at or below middle; leaf axis variously hairy, hairs to 1 mm, sometimes glandular; prickles generally ± thick-based, often curved; sepals often glandular; CA-FP, adjacent DMoj
12. Fls generally 1(6); hypanthium generally 5–7 mm wide at flower, (10)13–20 mm wide at fruit, generally ± spheric; achenes generally 4.5–6 mm — prickles to 20 mm, generally ± compressed, ± straight to ± curved; sepal tips generally > body, toothed ..... R. nutkana
13. Lflets generally ± single-toothed, generally glandless; prickles generally few; NCoRO, KR, CaRH; 750–1500 m ..... subsp. macdougalii
13' Lflets double-toothed, ± glandular especially beneath; prickles generally many; NCo, NCoRO, CCo; < 700 m ..... subsp. nutkana
12' Fls 1–many; hypanthium 2.5–5.5 mm wide at flower, 7–15(20) mm wide at fruit, spheric to (ob)ovoid; achenes 3–5 mm
14. Prickles generally thick-based, generally curved (straight), few to many; leaflets and leaf axis generally ± shaggy-hairy, hairs to 1 mm; pedicel generally ± hairy; hypanthium generally (ob)ovoid, glabrous to sparsely hairy, neck 2–4.5 mm wide at flower ..... R. californica
14' Prickles slender to ± thick-based, straight (± curved), 0–few; leaflets and leaf axis ± glabrous to ± hairy, hairs 0.1–1 mm; pedicels generally glabrous (except for glands); hypanthium spheric to ovoid, glabrous, neck ± 2 mm wide at flower ..... R. pisocarpa
15. Fls 1–3(10); sepals generally glandless; prickles 0 to paired at nodes; leaflets generally 5(7) ..... subsp. ahartii
15' Fls (1)3–10; sepals generally glandular; prickles generally paired at nodes; leaflets generally (5)7(9) ..... subsp. pisocarpa
Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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