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Key to families | Table of families and genera

This text currently parallels The Jepson Manual: Vascular Plants of California, Second Edition that is now available at the University of California Press.

Text appearing in blue on this page will not appear in the printed book; it will be displayed only on the Web. Specimen numbers are hyperlinked to records in the Consortium of California Herbaria data view where possible. Taxa are hyperlinked to entries in the Jepson Interchange via the "[Online Interchange]" link.

 

IVESIA

Barbara Ertter


Perennial, glandular; odor resinous.
Leaf: generally basal, odd-1-pinnately compound, generally ± cylindric; cauline generally alternate, reduced; leaflets 4–80 per side, generally overlapped, generally divided ± to base.
Inflorescence: cyme; pedicel bractlets 0.
Flower: receptacle generally not stalked; hypanthium shallow or deep, bractlets (0)5, generally < sepals; petals generally 5, 1–5(7) mm, linear to obovate or round, acute to rounded; stamens 5–20(40), filaments generally thread-like; pistils 1–8(20), ovary superior, style attached below fruit tip, base ± rough-thickened.
Fruit: achene.
30 species: w North America. (Eli Ives, Yale University, CT pharmacologist, 1779–1861) [Ertter & Reveal 2007 Novon 17:315–325] Lf, leaflet data for basal leaves.
Unabridged etymology: (Eli Ives, Yale University, CT pharmacologist, participant in Pacific Railroad Survey, 1779–1861)
Unabridged references: [Ertter 1989 Syst Bot 14:231–244]

Key to Ivesia

1. Lf ± flat, leaflets toothed or lobed generally < 3/4 to base, generally ± separate — plants rosettes or hanging clumps

2. Stamens 15–40 — s SNH, SnBr, PR, SNE, DMtns ..... I. saxosa

2' Stamens 5–10

3. Hypanthium bractlets 5; MP, adjacent SNH ..... I. baileyi

4. Petals pale yellow; hypanthium bractlets ± = sepals ..... var. baileyi

4' Petals white; hypanthium bractlets ± < 1/2 sepals ..... var. beneolens

3' Hypanthium bractlets generally 0; DMtns

5. Hypanthium width <= length, receptacle stalked in pistil-bearing portion; n DMtns ..... I. arizonica var. arizonica

5' Hypanthium width > 2 × length; receptacle not stalked; e DMtns ..... I. patellifera

1' Lf ± cylindric, leaflets generally lobed ± to base, overlapped

6. Stamens 10–20 (if 10, cauline leaves >= 3)

7. Lflets < 8 per side; stems hanging to ± matted; caudex few- to many-branched; rock outcrops

8. Lflet lobes < 4; petals white; SnJt ..... I. callida

8' Lflet lobes 3–6; petals yellow; e DMtns ..... I. jaegeri

7' Lflets > 10 per side; stems decumbent to erect; caudex 0–few-branched; meadows, sandy flats

9. Lflets generally 15–20 per side; sheathing leaf bases glabrous; stamens 10–16, filament ± < 1 mm; style < 1.5 mm

10. Petals 4(5), pale yellow; inflorescence not red-tinged; s SNH ..... I. campestris

10' Petals 5, white or pink-tinged; inflorescence often red-tinged; c&s SNH ..... I. unguiculata

9' Lflets >= 20 per side; sheathing leaf bases generally ± strigose; stamens 15–20; filament 1–4 mm; style ± > 2 mm

11. Infl of separate flowers or generally < 5-flowered clusters; pedicels generally > 5 mm; leaflets generally 30–80 per side

12. Lf mousetail-like, leaflets indistinct, obscured by dense, silvery hairs, lobes < 1.5 mm; stamens 15; pistil 1; granitic sand — SNH, TR, SnJt ..... I. santolinoides

12' Lf not mousetail-like, leaflets overlapped but distinct, not obscured by hairs, lobes >= 2 mm; stamens 20; pistils >= 2; meadows

13. Herbage hairs ± 0 or appressed, < 1 mm; petals ± obovate; alkali meadows; n SNE ..... I. kingii var. kingii

13' Herbage hairs spreading, < 4 mm; petals ± oblanceolate; generally on serpentine clays; c KR ..... I. pickeringii

11' Infl of generally > 5-flowered clusters; pedicels generally < 3 mm (except lowest); leaflets 20–35 per side

14. St 10–20 cm, ± decumbent; leaves generally 4–8 cm, cauline ± 2; filament ± flat; SnBr ..... I. argyrocoma var. argyrocoma

14' St generally > 20 cm, decumbent to erect; leaves > 8 cm, cauline >= 3; filament thread-like, not flat; n SNH, MP

15. Petals white; hypanthium length >= width; hairs of stem base ± spreading, 2–4 mm ..... I. sericoleuca

15' Petals yellow; hypanthium length <= width; hairs of stem base ascending, < 2 mm ..... I. aperta

16. Petals 2–3 mm, oblanceolate; filament 1–1.5 mm; n SNH (except Dog Valley), s MP ..... var. aperta

16' Petals generally 4–7 mm, ± obovate; filament 2–4 mm; n SNH (Dog Valley, e Sierra Co.) ..... var. canina

6' Stamens 5 or 10; cauline leaves 1–3

17. Hypanthium bractlets > sepals — KR (Castle Crags) ..... I. longibracteata

17' Hypanthium bractlets ± 1/2 sepals

18. Infl generally ± open between separate flowers or of ± loose clusters; plant ± matted from much-branched caudex; pedicels generally ± S-shaped in fruit; sheathing leaf bases strigose

19. Stamens 10; pistils 10–30 — c&s SNH ..... I. pygmaea (2)

19' Stamens 5; pistils generally < 5

20. Petals white to pale ± yellow, linear, ± 1 mm; plant densely hairy, glands not obvious; 1400–1800 m; MP ..... I. paniculata

20' Petals yellow, oblanceolate, ± 2 mm; plant moderately hairy, glands many; 2700–4000 m; n&c SNH, W&I ..... I. shockleyi var. shockleyi

18' Infl of 1–few generally head-like, 3–20-flowered clusters (becoming more open in Ivesia webberi); plant generally rosetted from 0–few-branched caudex; pedicels straight in fruit; sheathing leaf-bases hairy or not

21. Cauline leaves 2, ± opposite; leaflets 4–8 per side, lobes linear to lanceolate; 1500–1900 m — s-most MP, adjacent n SNH ..... I. webberi

21' Cauline leaves generally 1 or alternate; leaflets > 8 per side, lobes oblanceolate to ± round; > 1800 m

22. Lf mousetail-like, leaflets indistinct, < 1 mm, hidden by dense, silvery hairs; petals 1–2 mm — c&s SNH ..... I. muirii

22' Lf not mousetail-like, leaflets overlapped but distinct, generally > 1 mm, ± glabrous to ± hairy; petals >= 2 mm

23. Stamens 10; sheathing leaf-bases generally ± strigose; hypanthium length ± < l/2 width — c&s SNH ..... I. pygmaea (2)

23' Stamens 5; sheathing leaf-bases not strigose (± glabrous or glandular); hypanthium length > 1/2 width

24. Hypanthium length >= width; petals narrow-oblanceolate; pistils generally 2–4; fruit ± 2 mm, mottled brown ..... I. gordonii

25. Sts ascending to erect, generally not ± red ..... var. alpicola

25' Sts prostrate to ascending, dark ± red ..... var. ursinorum

24' Hypanthium length ± < width; petals ± obovate; pistils generally 5–15; fruit 1–1.5 mm, not mottled ..... I. lycopodioides

26. Lflet lobes ± 1 mm, ± round, ± glabrous; petals 2–3 mm — rocky areas; n&c SNH, SNE (Sweetwater Mtns) ..... var. lycopodioides

26' Lflet lobes >= 1 mm, narrow-oblanceolate to obovate, often ± hairy; petals generally > 3 mm

27. Lflet lobes 2–8 mm, ± glabrous to sparsely hairy, bristle-tip 0–0.5 mm; wet meadows; c&s SNH ..... var. megalopetala

27' Lflet lobes 1–3 mm, moderately to densely hairy, bristle-tip generally 0.5–1 mm; vernally moist, rocky areas; c&s SNH, W&I ..... var. scandularis

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