The Jepson Herbarium
		University of California, Berkeley

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Annual to shrub, matted or not.
Stem: prostrate to erect, occasionally 0, glabrous or hairy, occasionally glandular.
Leaf: basal, sheathing (on stem above basal, beneath other, ± non-sheathing cauline), or cauline, alternate, opposite, or whorled; blades linear to ± round, generally longer than wide, margins flat, wavy, or rolled under; ocreae 0.
Inflorescence: variable, glabrous or hairy, occasionally glandular; peduncles 0 or erect to reflexed (pointed down); involucres 1 or in clusters, tubular, glabrous or hairy, teeth generally 4–10, awns 0.
Flower: (2)6–100(200), with a stalk-like base (" flower stipe" or " stipe") or not; perianth generally white to red or yellow, glabrous or hairy, occasionally glandular, lobes 6, generally entire; stamens 9.
Fruit: generally brown or black, generally obconic, glabrous or hairy; embryo curved or straight.
± 250 species: temperate North America. (Greek: woolly knees, for hairy nodes of 1st sp. named) {hort link} One of largest genera in CA; stem in descriptions refers to the main stem(s), not branches of inflorescence. A per allied to Eriogonum austrinum (S. Stokes) Reveal and Eriogonum moranii Reveal of e-c Baja CA, with spreading (rather than reflexed or erect) involucres on peduncles < 2 mm, occurring in DMtns (Bristol, Granite mtns), has been known for nearly 25 yrs yet remains undescribed. Eriogonum puberulum moved to Johanneshowellia.

Key to Eriogonum

Key to Groups

1. Annual (except Eriogonum inflatum, perennial herb, keyed in Group 2 when flowering first yr)

2. Involucres angled to strongly ribbed, sessile and strongly appressed or at tips of short bracted branchlets (or peduncled) (subg. Oregonium) ..... Group 1

2' Involucres smooth, not ribbed or angled, generally peduncled, if sessile then not appressed (subg. Ganysma) ..... Group 2

1' Perennial

3. Fl stipe present, sometimes weakly so, not winged; inflorescence bracts 2–13+ (subg. Oligogonum) ..... Group 3

3' Fl stipe 0 or ± winged; inflorescence bracts generally 3 (mainly subg. Eucycla) ..... Group 4

Group 1: Ann; involucres angled to strongly ribbed, sessile and strongly appressed or at tips of short bracted branchlets (or peduncled) (subg. Oregonium)

1. Involucres at tips of slender branchlets, not appressed; peduncles present or 0

2. Fls 0.7–1.5 mm, yellow or cream; n SNE, w DMoj

3. Fls cream to ± red, 1–1.5 mm; inflorescence 5–10(15) cm wide; n SNE ..... E. ampullaceum

3' Fls yellow, 0.7–1 mm; inflorescence 5–20 cm wide; w DMoj ..... E. mohavense

2' Fls 1–2.5(3) mm, yellow or white to pink, rose, or red

4. St glabrous; involucres sessile

5. Basal leaf blades oblong-ovate; involucres 3–4 mm, 5-toothed; c NCoRI, n CCo, n SnFrB ..... E. luteolum var. caninum

5' Basal leaf blades generally round to reniform; involucres 2–4 mm, 5–8-toothed; c&s CW, WTR

6. Involucres 2–2.5 mm, 5-toothed; flowers 2–2.5(3) mm, minutely puberulent, white to rose or yellow in fruit; c&s CW, WTR ..... E. covilleanum

6' Involucres 3–4 mm, 6–8-toothed; flowers 1–2 mm, glabrous, white to rose; n SCoR ..... E. nortonii

4' St tomentose, if glabrous then involucres peduncled at lower nodes

7. Involucres glabrous; s SnFrB, n SCoRI ..... E. argillosum

7' Involucres tomentose; ScV, ne SnFrB, ne&s SCoRI

8. Lvs basal, blades ± round; styles 0.1–0.3 mm; s SCoRI ..... E. eastwoodianum

8' Lvs basal or basal and cauline, ± round or narrowly oblong to ovate; styles 0.2–1 mm; ne SnFrB, SCoRI

9. Involucres 2.5–3.5(4) mm; styles 0.2–0.3 mm; ScV (extirpated), ne SnFrB ..... E. truncatum

9' Involucres (1.5)1.8–2.5 mm; styles 0.2–1 mm; SCoRI

10. Fls smooth; involucres 2–2.5 mm; leaves basal and subbasal, rarely cauline; s SCoRI ..... E. temblorense

10' Fls papillate; involucres (1.5)1.8–2 mm; leaves basal and cauline; ne SCoRI ..... E. vestitum

1' Involucres not at tips of slender branchlets, generally appressed; peduncles 0

11. Fls hairy, white to rose — c&s NCoR ..... E. dasyanthemum

11' Fls glabrous or sparsely glandular, white, cream, yellow, rose, or red

12. Involucres 1–1.5 mm (except 2–4 mm in Eriogonum baileyi), if 2 mm then outer perianth lobes fan-shaped or ± hastate

13. Sts glabrous, if tomentose then flowers glandular and plant of e SN, GB; outer perianth lobes oblong to oblong-obovate

14. Fls yellow or pale ± yellow (or ± white), 0.6–0.8(1) mm; SNE, DMoj ..... E. brachyanthum

14' Fls white to rose, 1–2 mm; SNF, Teh, SnFrB, s SCoR, TR, GB, w DMoj

15. Fls 1–1.5 mm, glabrous or puberulent; stem, inflorescence glabrous — SnFrB, SCoR, WTR ..... E. elegans

15' Fls 1.5–3 mm, generally minutely glandular, occasionally glabrous in SNF; stem, inflorescence glabrous or tomentose ..... E. baileyi

16. Infl glabrous; SNF, Teh, s SCoR, TR, GB, w DMoj ..... var. baileyi

16' Infl tomentose; GB ..... var. praebens

13' Sts tomentose to sparsely so; outer perianth lobes narrowly to widely fan-shaped or ± hastate

17. Outer perianth lobes ± hastate in fruit; flowers 0.8–1.2 mm; involucres bell-shaped, 1–2 mm; SnBr, PR ..... E. evanidum

17' Outer perianth lobes fan-shaped; flowers 1–3.5 mm; involucres bell-shaped to obconic, 1–2.5 mm; GB, D

18. Lvs cauline; leaf blades narrowly oblanceolate to widely elliptic; plant narrowly erect; flowers white to pink or red; se DMoj ..... [E. polycladon]

18' Lvs basal; leaf blades ± round to cordate; plant generally spreading; flowers white to pink or pale yellow to yellow; GB, D

19. Fls pale yellow or yellow to red; outer perianth lobes widely fan-shaped; inflorescence generally densely branched, branches incurved; fruit 1–1.3 mm; GB, D ..... E. nidularium

19' Fls white to pink or pale yellow, outer perianth lobes narrowly fan-shaped; inflorescence generally sparsely branched, branches not incurved; fruit 1.5–1.8 mm; SNE, D ..... E. palmerianum

12' Involucres (1.8)2–5(7) mm; outer perianth lobes not fan-shaped or hastate (see also Eriogonum baileyi at 12.)

20. Lf blades oblong-obovate to oblanceolate or oblong (basal occasionally ± rounded in E. cithariforme)

21. Involucres (3.5)4–5 mm; outer perianth lobes narrowly obovate to oblong; fruit 1.8–2(2.2) mm; NW, SNF, Teh, CW, TR ..... E. roseum

21' Involucres (1.8)2–3 mm; outer perianth lobes lanceolate to oblong or oblong-obovate; fruit 1–2 mm; CA-FP

22. Infl branches upcurved; basal leaf blades oblanceolate or elliptic to ovate or ± rounded, (0.3)1–2 cm; petiole generally winged; involucres 2.5–3 mm; flowers 1.5–2 mm, white to rose; SCoR, TR ..... E. cithariforme

23. St, inflorescence glabrous; SCoR, WTR ..... var. agninum

23' St, inflorescence tomentose; SCoR, TR ..... var. cithariforme

22' Infl branches straight; basal leaf blades oblanceolate to oblong, (0.8)1–4(6) cm; petiole not winged; involucres (1.8)2–3 mm; flowers 1.5–3 mm, white to pink or yellow; CA-FP ..... E. gracile

24. Infl densely to sparsely tomentose; widespread ..... var. gracile

24' Infl glabrous; PR ..... var. incultum

20' Lf blades round-ovate to rounded or reniform, if not then stem and inflorescence branches glabrous and plant of SNH; stem, inflorescence branches glabrous to tomentose

25. Infl sparsely tomentose, especially basally, or glabrous; leaves basal; flowers white to rose or pale yellow — ne KR, CaR, n SNH, MP ..... E. vimineum

25' Infl tomentose or generally glabrous, if sparsely tomentose then leaves basal and cauline; flowers white to pink, red or yellow

26. Involucres (3.5)4–5(7) mm; plant 4–10 dm; n WTR, SnBr, PR ..... E. molestum

26' Involucres 2–4 mm; plant 1–5 dm; widespread

27. Lvs basal; involucres 3–4 mm; flowers 1.5–2 mm; stem glabrous; SW, SNE, DMoj ..... E. davidsonii

27' Lvs basal and cauline, involucres 3–3.5 mm and flowers 1.5–2 mm with sparsely tomentose stem and inflorescence, if leaves only basal and stem glabrous then involucres 2–3.5 mm and plant of n&c SN; n CA-FP ..... E. luteolum (in part)

28. Involucres 2–3 mm; flowers 2–2.5 mm; plant of granitic sand; n&c SNH ..... var. saltuarium

28' Involucres 3–3.5 mm; flowers 1–2 mm; plant of serpentine; NW, CaR, SNF, SnFrB

29. Fls 1.8–2 mm; NW, CaR, SnFrB ..... var. luteolum

29' Fls 1–1.5(1.8) mm; SNF ..... var. pedunculatum

Group 2: Ann (exc 1st-yr fl Eriogonum inflatum); involucres smooth, not ribbed or angled, gen peduncled, if sessile then not appressed (subg. Ganysma)

1. Perennial, flowering first yr or not — GB, D ..... E. inflatum (2)

1' Annual (per Eriogonum inflatum may key here if flowering first yr)

2. Lvs basal and cauline, not basal only or basal and sheathing

3. Involucres glandular-hairy or puberulent

4. Perianth lobes alike, oblong to elliptic, not inflated

5. Involucre glandular-hairy, densely tomentose adaxially, 2.7–3 mm; flower 1.5–1.7 mm; s SNF, sw SnJV ..... E. gossypinum

5' Involucre not densely tomentose, 1.8–2 mm; flower 2–2.5 mm; s SN, SCoR, TR ..... E. gracillimum

4' Perianth lobes not alike, outer elliptic to obovate or ± round, inflated or not, inner narrower

6. Outer perianth lobes generally elliptic to obovate, not obviously inflated, if so then only basally; stamens exserted; c&s SNF, Teh, SnJV, CW, TR, s PR, w DMoj ..... E. angulosum

6' Outer perianth lobes elliptic to ± round or obovate, obviously inflated basally or at tip; stamens included; SnJV, SCoR, s SN, TR, GB, w DMoj

7. Outer perianth lobes inflated base to middle; involucres glandular-hairy; s SN, TR, GB ..... E. maculatum

7' Outer perianth lobes inflated above middle; involucres glandular; SnJV, SCoR, TR, w DMoj ..... E. viridescens

3' Involucres glabrous, long-stiff or long-soft hairy, not glandular-hairy or puberulent

8. Fls hairy, hairs hooked

9. Fls 2, 0.8–1.1 mm; fruit tip exserted; NW, SN, n CW, TR ..... E. hirtiflorum

9' Fls 4–6, 1.2–1.8 mm; fruit tip not exserted; s NCoR, c&s SN, Teh, CW, WTR ..... E. inerme

10. Involucres short-stiff hairy; c&s SN, Teh ..... var. hispidulum

10' Involucres glabrous; s NCoR, CW, WTR ..... var. inerme

8' Fls glabrous, if hairy then hairs not hooked

11. Lvs sparsely tomentose or glabrous, oblong-oblanceolate to obovate; flowers densely short-hairy; plant glabrous or sparsely tomentose; s SNF, Teh, SCoRI, n WTR ..... E. ordii (2)

11' Lvs short-hairy, linear; flowers glabrous or sparsely hairy; plant glabrous or glandular and short-hairy; NCoRH, CaRH, SN, n SCoRO, TR ..... E. spergulinum

12. Infl internodes not glandular; plant prostrate to ascending — s SNH ..... var. pratense

12' Infl internodes glandular; plant erect

13. Fl 1.5–2.5 mm; NCoRH, CaRH, SN, n SCoRO, TR ..... var. reddingianum

13' Fl 2.5–3.5 mm; SN ..... var. spergulinum

2' Lvs basal and sometimes sheathing, not both basal and cauline

14. Lf blades glabrous, or hairy on one or both surfaces, or if sparsely tomentose then flowers densely short-soft-hairy

15. Fls generally glabrous — c SNH, SNE ..... E. esmeraldense var. esmeraldense

15' Fls hairy

16. Fls densely short-soft hairy or puberulent, white to pink or red (or ± yellow)

17. Lf blades oblong-oblanceolate to obovate, sparsely tomentose or glabrous; stem glabrous to sparsely tomentose; s SNF, Teh, SCoRI, n WTR ..... E. ordii (2)

17' Lf blades oblanceolate to obovate, sparsely soft-shaggy hairy and glandular or coarse-hairy; stem glabrous and sparsely glandular; s SN, TR, PR, n W&I, sw DMtns

18. Fls pink to red, aging to white, 0.5–0.9 mm; leaf blades coarse-hairy; involucres 0.5–0.9 mm; s SN, TR, PR, n W&I ..... E. parishii

18' Fls white to ± red (or yellow), 1.5–2.5 mm; leaf blades soft-shaggy hairy, some glandular; involucres 1.2–1.5 mm; e SnBr, c&s PR, sw DMtns ..... E. apiculatum

16' Fls soft-shaggy- or coarse-hairy, white to ± pink or generally yellow

19. Plant glandular throughout; flowers white to ± pink — n&ne DMtns ..... E. glandulosum

19' Plant glabrous, except sometimes glandular or hairy near base or at nodes; flowers yellow to green-yellow or ± red

20. St glandular near base

21. Infl glandular at lower nodes; involucre teeth (4)5; plant (0.3)0.5–3 dm; leaf blades generally ± round, (0.3)0.5–1(1.4) cm; fruit 1.5–1.8(2) mm; ne DMoj ..... E. contiguum

21' Infl glabrous; involucre teeth 4; plant (2)4–18(22) dm; leaf blades round to reniform, (0.5)1–2.5(4) cm, (0.5)1–2(3) cm wide; fruit 2–2.5 mm; CW, SW, s SNE, D ..... E. clavatum (2)

20' St glabrous or basally coarse- or long-soft-hairy

22. St, inflorescence ± gray; perennial herb but appearing annual when flowering first yr — GB, D ..... E. inflatum (2)

22' St, inflorescence branches ± green or yellow-green; annual

23. Involucres 1–1.5(1.8) mm; plant (2)4–18(22) dm; inflorescence 30–150(170) cm; fruit 2–2.5 mm; CW, SW, s SNE, D ..... E. clavatum (2)

23' Involucres 0.7–1 mm; plant 1–4.5(6) dm; inflorescence 5–30 cm; fruit 1–1.5 mm; D ..... E. trichopes

14' Lf blades tomentose to sparsely so on one or both surfaces; flowers glabrous, glandular, or variously hairy, but not short-soft-hairy

24. Outer perianth lobes generally oblong, obovate or round, cordate basally

25. Involucres sessile and erect or on erect peduncles <= 0.5 cm — DMoj

26. St 0.5–3 cm; plant 1–4 dm; inflorescence branches spreading ..... E. bifurcatum

26' St (5)10–20 cm; plant (0.5)3–6(10) dm; inflorescence branches ± erect ..... E. exaltatum

25' Involucres sessile and reflexed or on reflexed peduncles <= 1.5 cm

27. St, inflorescence glandular; SNE, DMoj ..... E. brachypodum

27' St, inflorescence glabrous; s SNH, Teh, s SnJV, TR, GB, D

28. Involucres widely bell-shaped; peduncles 0; flowers yellow to red-yellow; SNE ..... E. hookeri

28' Involucres narrowly obconic or narrowly bell-shaped; peduncles <= 1.5 cm; flowers white to pink; s SNH, Teh, s SnJV, TR, D

29. Involucres 1–1.5 mm; inflorescence of many horizontal tiers; ne DMoj ..... E. rixfordii

29' Involucres 1.5–2.5 mm; inflorescence not of many horizontal tiers; s SNH, Teh, s SnJV, TR, D ..... E. deflexum (in part)

30. Involucres (2)2.5–3 mm; peduncles (0.3)0.5–1.5 cm; stem inflated; s SNH, Teh, s SnJV, TR, n&w DMoj ..... var. baratum

30' Involucres 1.5–2.5 mm; peduncles 0 or generally <= 0.5 cm; stem not inflated; D ..... var. deflexum

24' Outer perianth lobes oblong to oblanceolate or ovate, truncate to obtuse basally

31. Fls papillate basally — peduncles curving, ascending, 1–5 cm; MP ..... E. collinum

31' Fls smooth, not papillate

32. Perianth lobes alike

33. Sts, inflorescence glandular; peduncles 0 or reflexed; flowers 2–2.5 mm ..... E. eremicola

33' Sts, inflorescence glabrous; peduncles erect; flowers 1.5–1.8 mm ..... E. hoffmannii

34. Lf margins flat, blades 2–4 cm wide; petioles 1–5 cm; plant 0.5–5 dm ..... var. hoffmannii

34' Lf margins wavy, blades 3–8 cm wide; petioles (3)5–10 cm; plant 4–10 dm ..... var. robustius

32' Perianth lobes not alike, if alike then glandular-hairy

35. Fls glandular

36. Fls white to red, glandular-hairy with a tuft of long white hairs adaxially; e PR, s DMoj, DSon; outer perianth lobes generally fan-shaped ..... E. thurberi

36' Fls yellow, if in age ± white then outer perianth lobes sac-like basally, glandular or short-stiff hairy, without a tuft of long white hairs adaxially; SCoRI, WTR, GB, D

37. Outer perianth lobes cordate, in age sac-like basally, initially yellow then white to rose; involucres 0.6–1.2 mm, glabrous; D ..... E. thomasii

37' Outer perianth lobes oblong-elliptic to obovate or ovate, not sac-like basally, yellow; involucres 1–2 mm, glabrous or glandular-hairy; SCoRI, WTR, GB, D

38. Involucres glandular-hairy; SCoRI, WTR, GB, D ..... E. pusillum

38' Involucres glabrous; D ..... E. reniforme

35' Fls glabrous

39. Peduncles 0 or reflexed; outer perianth lobes oblong; SNE ..... E. deflexum var. nevadense

39' Peduncles generally curving downward; outer perianth lobes ± fiddle-shaped or oblong to oval; n&c SNH, GB

40. Outer perianth lobes ± fiddle-shaped; peduncles glabrous (or 0); c SNH, GB, probably introduced in n SNH (Nevada Co.) ..... E. cernuum

40' Outer perianth lobes oblong to oval; peduncles glandular (glabrous); n SNH, n SNE ..... E. nutans

41. Peduncles glandular; n SNE ..... var. nutans

41' Peduncles glabrous; n SNH (introduced but now extirpated) ..... [var. glabratum]]

Group 3: Per; fl stipe present, sometimes weakly so, not winged; infl bracts 2–13+ (subg. Oligogonum)

1. Involucre teeth not lobe-like, << tube, erect or ± so

2. Fls hairy

3. Lf blades tomentose to densely so abaxially, generally glabrous adaxially; nw KR ..... E. pyrolifolium var. coryphaeum

3' Lf blades glabrous or short hairy on both surfaces; n KR, CaRH, c&s SNH, W&I, DMtns

4. Lf blades glabrous; flowers white to rose; stem generally ascending; CaRH ..... E. pyrolifolium var. pyrolifolium

4' Lf blades short hairy; flowers cream to yellow; stem erect; n KR, W&I, DMtns

5. Fls bright yellow, 3–3.5(4) mm; n KR ..... E. hirtellum

5' Fls cream to pale yellow, 3–6 mm; c&s SNH, W&I, n DMtns ..... E. latens

2' Fls glabrous

6. Infl not immediately subtended by a whorl of bracts; stem with whorl of bracts near middle

7. Fls ± white to pink or rose-red; leaf blades silky-tomentose abaxially, 0.1–0.3(0.4) cm wide; c NCoRO ..... E. kelloggii

7' Fls yellow; leaf blades tomentose abaxially, 0.2–2(3) cm wide; not c NCoRO

8. Sts (0.3)0.4–0.6 dm; leaf blades 1–2(3) cm wide — e KR ..... E. alpinum

8' Sts 0.5–3 dm; leaf blades 0.2–0.7(0.8) cm wide

9. Involucre teeth 5–8, 0.5–1.5 mm; s KR, n NCoRH, n NCoRI ..... E. libertini

9' Involucre teeth 8–10, 1–3 mm; SN (see also 29') ..... E. prattenianum (2)

6' Infl immediately subtended by whorl of bracts; stem without a whorl of bracts near middle

10. Fls chalky-white, white to rose or ± pale yellow; flower stipe 0.1–0.4 mm

11. Fls white to rose, 5–7 mm; involucres 5–10(12) mm; NW, CaR, SN, GB ..... E. lobbii (2)

11' Fls chalky-white, 2.5–3.5 mm; involucres (2)2.5–3.5(4) mm; s SNH ..... E. polypodum

10' Fls yellow or ± brown to sulphur yellow or cream, occasionally with blush of pink-red to maroon (or white); flower stipe (0.1)0.3–1.5 mm

12. Plant unisexual, pistillate and staminate plants morphologically different

13. Lf blades ± glabrous, bright green to olive-green adaxially; e KR, CaRH, n&c SNH ..... E. marifolium

14. Involucres 4–5 mm, 5–7 mm wide; leaf-blades narrow; CaRH ..... var. cupulatum

14' Involucres 2–3 mm, 1.5–2 mm wide; leaf-blades wide to round; e KR, CaRH, n&c SNH ..... var. marifolium

13' Lf blades tomentose to densely so, generally not green adaxially; KR, SNH

15. Pistillate inflorescence generally persistently head-like (or umbel-like after fertilization); leaf blades (0.5)1–2 cm, 0.5–1.5 cm wide, densely tomentose; petioles 0.7–3 cm; KR ..... E. diclinum

15' Pistillate inflorescence initially head-like, umbel-like after fertilization; leaf blades 0.5–1.5 cm, 0.3–0.7 cm wide, generally tomentose; petioles (0.3)0.5–1 cm; SNH ..... E. incanum

12' Plant bisexual (but generally with some aborted male or female parts), otherwise not morphologically different

16. Fls sulphur yellow; involucres long-soft-wavy-hairy; inflorescence often compound umbel-like

17. Lf blades narrow-elliptic to -oblong, 0.3–0.6(0.8) cm wide; subshrub; KR, CaRH ..... E. congdonii

17' Lf blades oblong to obovate, (0.4)0.8–1.3 cm wide; perennial herb mat; n KR, s NCoRO, n NCoRH ..... E. ternatum

16' Fls white, cream, pale yellow (yellow); involucres tomentose; inflorescence umbel-like

18. Sts 0.2–0.6(1) dm; inflorescence ± head-like; flowers yellow to red or maroon, or white to pink-rose or deep red; s NCoRH, c&s NCoRI

19. Involucres 4–6.5 mm; stipe 0.1–0.3 mm; flowers yellow to red or maroon; s NCoRO ..... E. cedrorum

19' Involucres 3–4 mm; stipe 0.5–0.8 mm; flowers white to pink-rose or deep red; s NCoRH, c&s NCoRI ..... E. nervulosum

18' Sts 0.4–4 dm; inflorescence compound-umbel- or umbel-like; flowers cream or ± yellow, blushed or not with pink-red to maroon; KR, s CaR, n SN ..... E. ursinum

20. Fls 5–9 mm, cream (or yellow); fruit (5)5.5–8 mm; KR ..... var. erubescens

20' Fls 4–6 mm, pale yellow (or yellow); fruit 3–3.5 mm; s CaR, n SN ..... var. ursinum

1' Involucre teeth lobe-like, at least 1/2 tube, generally reflexed or spreading

21. Fls hairy

22. St bractless — GB ..... E. caespitosum

22' St with whorl of subtending bracts at base of umbel, middle of stem, or middle of branches

23. Infl scapose, involucres 1

24. Lf blades densely tomentose on both surfaces; e KR, e CaRH, n SNH, MP ..... E. douglasii var. meridionale

24' Lf blades sparsely tomentose to ± glabrous adaxially; CaRH, n&s SNH, MP

25. Lf blades generally narrowly oblanceolate; CaRH, n SNH, MP ..... E. sphaerocephalum var. halimioides (2)

25' Lf blades oblanceolate to elliptic; s SNH ..... E. twisselmannii

23' Infl not scapose, involucres >= 1

26. Fls 4–5 mm, yellow; leaf blades densely tomentose; plant 2.5–5 dm; NCoRI, n&c SNF ..... E. tripodum

26' Fls (5)6–9 mm, cream or pale to bright yellow; leaf blades densely tomentose to glabrous; plant 0.5–4 dm; CaRH, n SNH, MP ..... E. sphaerocephalum

27. Fls pale yellow to cream; CaRH, n SNH, MP ..... var. halimioides (2)

27' Fls bright yellow; n SNH, MP ..... var. sphaerocephalum

21' Fls glabrous

28. Infl head-like, involucres not immediately subtended by leafy bracts; stem with a whorl of leafy bracts near middle; flowers yellow

29. Lf blades (0.3)0.5–0.8 cm, sparsely tomentose to ± glabrous, bright green or olive-green adaxially; generally matted per; e KR ..... E. siskiyouense

29' Lf blades 0.5–1.5(2) cm, densely tomentose or ± glabrous adaxially, generally pale green adaxially; plant erect or rounded per or subshrub, occasionally mat; n&c SN, s SNH ..... E. prattenianum (2)

30. Lf blades tomentose abaxially, subglabrous adaxially; c&s SNH ..... var. avium

30' Lf blades tomentose on both surfaces; n&c SN ..... var. prattenianum

28' Infl umbel-like or compound-umbel-like, involucres immediately subtended by leafy bracts; stem with or without an additional whorl of bracts near middle; flowers white, yellow, or rose

31. St with whorl of leaf-like bracts near middle — leaf blades generally linear-oblanceolate to oblanceolate; Wrn ..... E. heracleoides var. heracleoides

31' Sts without a whorl of leaf-like bracts near middle

32. Fl stipe 0.1–0.4 mm — stem prostrate to decumbent or weakly erect; inflorescence ± head- to umbel-like — NW, CaR, SN, GB ..... E. lobbii (2)

32' Fl stipe 0.7–2 mm

33. Lf blades (2)7–25 cm; stem generally ± inflated; NW, CaR ..... E. compositum var. compositum

33' Lf blades 0.3–3(4) cm; stem not inflated; most of CA ..... E. umbellatum

34. Main inflorescence branches of 1 order, none with a whorl of bracts near middle

35. Fls generally ± white or cream to red, occasionally ± yellow

36. Lf blades densely tomentose on both surfaces — flowers generally lemon yellow to yellow-red; SnGb, SnBr ..... var. minus

36' Lf blades tomentose to glabrous adaxially

37. Fls pale yellow to cream or ± white (or green-white); leaf blades ± green adaxially; Wrn, W&I ..... var. dichrocephalum

37' Fls yellow, in age red-brown to rose or pink, with large ± red spot on each lobe; leaf blades generally ± red adaxially; W&I (Inyo Mtns), DMtns ..... var. versicolor (2)

35' Fls bright yellow

38. Fl clusters with branches generally > 2.5 cm

39. Lf blades sparsely tomentose abaxially; subshrub — SN, GB, nw DMoj ..... var. nevadense (2)

39' Lf blades densely white-tomentose or tomentose abaxially; subshrub or spreading to prostrate mat

40. Shrub 3–5 dm, on non-serpentine; KR, NCoRH, NCoRI, CaR, n SN, MP ..... var. dumosum

40' Perennial mat (0.7)1–4.5(5) dm, on serpentine; KR, n NCoRH, n CaRH

41. Sts 1–2.5(4) dm; leaf blades 0.5–2(3.5) cm; flowers 6–8(9) mm ..... var. goodmanii

41' Sts 0.5–1.5 dm; leaf blades 0.5–1(1.5) cm; flowers 3–6 mm ..... var. humistratum

38' Fl clusters with branches <= 2.5 cm

42. Plant prostrate, subalpine and alpine — c&s SNH, n W&I ..... var. covillei

42' Plant erect to ± spreading, generally not subalpine or alpine

43. Lf blades sparsely tomentose abaxially; SN, GB, nw DMoj ..... var. nevadense (2)

43' Lf blades tomentose to densely so abaxially; KR, NCoRO, CaR, SN, MP, DMtns

44. Infl branches (2.5)3–10(15) cm; CaR, n SN, MP ..... var. modocense

44' Infl branches 1–2.5(4) cm; KR, s SN, DMtns

45. Involucre tubes 3–4 mm; plant on serpentine; KR ..... var. nelsoniorum

45' Involucre tubes 2–3 mm; plant not on serpentine; s SNH, n DMtns ..... var. canifolium

34' Main inflorescence branches of >= 2 orders, or at least some with a whorl of bracts near middle

46. Infl with a whorl of bracts at middle (see also Eriogonum umbellatum var. polyanthum)

47. Fls cream or ± white, 4–7 mm; Wrn ..... var. glaberrimum

47' Fls yellow, 7–10 mm; n SNH ..... var. torreyanum

46' Infl without a whorl of bracts at middle, bracts only at base of inflorescence or involucres

48. Fls 7–10(12) mm; shrub; inflorescence 2–4-branched; KR ..... var. speciosum

48' Fls 3–8 mm; per mat or subshrub, if shrub then inflorescence 3-branched; not of KR

49. Fls cream, ± white, pale yellow, or green-yellow, or yellow, in age red-brown to rose or pink

50. Fls cream, ± white, pale yellow, or green-yellow, without large ± red spot on each lobe; subshrub or shrub; e DMtns ..... var. juniporinum

50' Fls yellow, in age red-brown to rose or pink, with large ± red spot on each lobe; spreading to ± prostrate mat; W&I (Inyo Mtns), DMtns ..... var. versicolor (2)

49' Fls yellow, in age not red-brown to rose or pink

51. Lf blades glabrous to densely tomentose (see also Eriogonum umbellatum var. munzii of TR, SnJt)

52. Lf blades densely tomentose — NCoRI, CW ..... var. bahiiforme

52' Lf blades densely tomentose to glabrous

53. Lf blades tomentose to densely so abaxially, tomentose to sparsely so adaxially; NW

54. Lf blades (0.5)0.8–1.8(2) cm wide; involucre lobes (3)4–6 mm; KR ..... var. lautum (2)

54' Lf blades 0.3–0.7 cm wide; involucre lobes 1–3 mm; NCoR ..... var. smallianum (2)

53' Lf blades generally sparsely tomentose to glabrous; not NW

55. Lf blades glabrous; s SNH, SNE, nw DMoj ..... var. chlorothamnus

55' Lf blades sparsely tomentose to glabrous adaxially; se SN, TR, SNE, DMtns ..... var. subaridum

51' Lf blades sparsely to densely tomentose, or densely tomentose abaxially, less so to glabrous adaxially (rarely both surfaces tomentose in Eriogonum umbellatum var. munzii of TR, SnJt)

56. Shrub, densely branched — n SNF, n SNH/CaRH

57. Lf blades rusty-woolly to tomentose abaxially; inflorescence 3–4-branched, branches tomentose to woolly, central branch without a whorl of bracts near middle ..... var. ahartii

57' Lf blades white-tomentose abaxially; inflorescence 1–2(3)-branched, branches sparsely tomentose to glabrous, central branch occasionally with a whorl of bracts near middle ..... var. polyanthum

56' Perennial mat or more openly and sparsely branched subshrub or shrub

58. Sts generally sparsely tomentose or glabrous

59. Lf margins finely wavy; stems often with leaf-like bract near middle; KR, c NCoRH ..... var. argus

59' Lf margins flat; stems without leaf-like bract near middle; SNH ..... var. furcosum

58' Sts generally tomentose or sparsely tomentose (or ± glabrous in age)

60. Involucre lobes (3)4–6 mm; leaf blades generally widely elliptic — KR ..... var. lautum (2)

60' Involucre lobes 1–3 mm; leaf blades elliptic

61. Lf blades 0.5–1 cm wide; TR, SnJt ..... var. munzii

61' Lf blades 0.3–0.7 cm wide; NCoR ..... var. smallianum (2)

Group 4: Per; fl stipe 0 or ± winged; infl bracts gen 3 (mainly subg. Eucycla)

1. St basally jointed, breaking into short-cylindric sections; flowers 150–200 — n DMtns (subg. Clastomyelon) ..... E. intrafractum

1' St not jointed, not breaking into sections; flowers 7–100

2. Fls with dense coarse, curved, ± white hairs; leaves generally coarse-hairy on both surfaces; GB, D (subg. Ganysma) ..... E. inflatum

2' Fls glabrous or not as above; leaves tomentose to sparsely so at least abaxially, not coarse-hairy

3. Fl stipe ± winged

4. Fls bright yellow; inflorescence cyme- or umbel-like, 0.5–3 cm; fruit 2.5–3 mm; s WTR ..... E. crocatum

4' Fls white to rose or ± yellow; inflorescence cyme-like, 10–25 cm; fruit 3.5–4 mm; s SN, CW, SW, SNE, DMtns ..... E. saxatile

3' Fl stipe 0

5. Plant forming cespitose or cushion-like mat; inflorescence head-like, if not then plant 0.1–1 dm

6. Infl branched

7. Fls glandular-hairy, ± white to ± red; stem generally glandular-hairy; s SNH ..... E. breedlovei

8. St erect; inflorescence umbel-like (head-like) ..... var. breedlovei

8' St suberect to prostrate; inflorescence cyme-like ..... var. shevockii

7' Fls glabrous, white to pink or rose or ± yellow with ± red spot; stem not glandular; s SNH, n DMtns

9. Fls 3.5–5 mm, lobes not alike, outer round; involucres 1.5–2 mm; n DMtns ..... E. gilmanii

9' Fls (1.5)2–2.5 mm, lobes alike; involucres 0.8–1.7(2) mm; s SNH ..... E. wrightii var. olanchense (2)

6' Infl head-like

10. Perianth lobes not alike, outer generally 2 × as wide as inner; stems sparsely to densely tomentose or ± glabrous — KR, CaR, SNH, SnBr, GB ..... E. ovalifolium

11. Lf blades generally 1–6 cm (or shorter); stem (1)5–30(40) cm; involucres (3.5)4–6.5(8) mm

12. St generally 1–5(7.5) cm; — leaf blades densely tomentose; n SNH (Job's Peak, Alpine Co.) ..... var. eximium (2)

12' St generally (4)7–30(40) cm (or shorter)

13. Fls 5–7 mm; involucres 5–7 mm; rare, ne SnBr ..... var. vineum (2)

13' Fls 4–5 mm; involucres 4–6.5 mm; common, not SnBr

14. Fls yellow; SNH, GB ..... var. ovalifolium

14' Fls white to rose or purple; KR, CaRH, SNH, GB ..... var. purpureum

11' Lf blades 0.2–1.2 (2) cm; stem generally 0.3–5(9) cm (or longer); involucres generally 2–4.5(8) mm

15. Fls yellow — c SNH ..... var. caelestinum

15' Fls white, rose, purple or red

16. Lf blades densely tomentose, margins ± brown — n SNH (Job's Peak, Alpine Co.) ..... var. eximium (2)

16' Lf blades tomentose to densely so, margins not ± brown

17. Involucres 3–4.5 mm

18. Lf blades tomentose, to sparsely so adaxially; rare, MP ..... var. depressum

18' Lf blades tomentose to densely so on both surfaces; common, CaRH, SNH, W&I ..... var. nivale

17' Involucres 5–8 mm — s SNH or ne SnBr

19. Lf blades tomentose to sparsely so; s SNH ..... var. monarchense

19' Lf blades densely tomentose; ne SnBr ..... var. vineum (2)

10' Perianth lobes alike, if not then stems glandular-hairy

20. Fls yellow or red-yellow

21. Involucres glandular and sparsely hairy; stem glandular-hairy; n&c SNH, n SNE ..... E. rosense var. rosense

21' Involucres glabrous or sparsely tomentose, occasionally glandular; stem glabrous or glandular; MP

22. Lf blades obovate or not, 1–2 cm, (0.3)0.5–1(1.5) cm wide; stem 0.5–1.5(2) dm, glandular below inflorescence; involucres sparsely tomentose, occasionally sparsely glandular ..... E. ochrocephalum var. ochrocephalum

22' Lf blades not obovate, 0.3–1(1.4) cm, 0.15–0.4(0.6) cm wide; stem 0.2–0.6(0.8) dm, glabrous; involucres glabrous or sparsely tomentose ..... E. prociduum

20' Fls white to rose or pink

23. Involucres sparsely glandular to glandular-hairy or hairy, pliable; c&s SNH, n W&I ..... E. gracilipes

23' Involucres glabrous or tomentose, rigid; c&s SN, Teh, TR, SNE, n DMtns

24. Fr tomentose; SNE, n DMtns (Last Chance Range) ..... E. shockleyi var. shockleyi

24' Fr glabrous; c&s SN, Teh, TR, SNE, nw DMtns

25. Pedicels hairy; leaves (1)2–5 cm, 0.8–2 cm wide; plant 3–10(11) dm diam; Teh ..... E. callistum (2)

25' Pedicels glabrous; leaves 0.2–1(1.2) cm, 0.05–0.4 cm wide; plant 1–4 dm diam; c&s SN, TR, SNE, nw DMtns ..... E. kennedyi

26. Lf hairs white; plant forming dense mat; c&s SN, SNE, nw DMtns ..... var. purpusii

26' Lf hairs gray- or red- to brown-white; plant generally forming loose, open mat; se SNF, TR

27. Lf blades oblong, 0.3–0.5 cm, 0.1–0.4 cm wide; se SNF ..... var. pinicola

27' Lf blades oblanceolate to elliptic, 0.2–1.2 cm, 0.05–0.2 cm wide; TR

28. St 0.5–2(3) cm; SnGb, SnBr ..... var. alpigenum

28' St 4–15 cm; n WTR, SnBr

29. St sparsely tomentose; involucres 2.5–4 mm ..... var. austromontanum

29' St glabrous; involucres 1.5–2.5 mm ..... var. kennedyi

5' Shrub to subshrub or spreading to erect herb; inflorescence branched, if head-like then plant 1–5 dm

30. Involucres clustered 2–10+ per node

31. Perianth lobes not alike ..... E. strictum

32. Fls yellow — CaR, ne SN, MP ..... var. anserinum

32' Fls white to rose or purple

33. Lf hairs white, densely tomentose to tomentose; KR ..... var. greenei

33' Lf hairs ± gray or ± green, tomentose to sparsely so; KR, n NCoRO, NCoRH, CaR, ne SNH, w MP ..... var. proliferum

31' Perianth lobes alike

34. Herb

35. Lf blades lanceolate to narrowly ovate, 4–15(25) cm, long-soft-hairy ..... E. elatum

36. St, inflorescence glabrous; n CA ..... var. elatum

36' St, inflorescence long-soft-hairy; KR, CaR, n SN, MP ..... var. villosum

35' Lf blades oblanceolate to ovate or elliptic, generally 2–6 cm, tomentose abaxially, tomentose to glabrous adaxially

37. Involucres 3–6 mm wide; stem, inflorescence glabrous (to sparsely tomentose); inflorescence ± head- to cyme-like; leaf blades densely tomentose abaxially, tomentose to ± glabrous adaxially; ChI ..... E. grande

38. Fls pink to red or rose; plant 2–5 dm; involucres 5–7 mm; Santa Rosa, San Miguel, Santa Cruz islands, occasionally CW ..... var. rubescens

38' Fls white; plant 1–15 dm; involucres 4–6 mm; various islands except Santa Rosa Island

39. Plant 5–15 dm; involucres 5–6 mm; San Miguel, Santa Cruz, Anacapa, Santa Catalina, San Clemente islands, occasionally CW ..... var. grande

39' Plant 1–2(2.5) dm; involucres 4–5 mm; San Nicolas Island ..... var. timorum

37' Involucres (1.5)2–4 mm wide; stem, inflorescence glabrous to tomentose; inflorescence head- to umbel- or cyme-like; leaf blades tomentose to densely so on both surfaces, or tomentose to glabrous adaxially; mainland

40. Pedicels hairy; flowers densely hairy — Teh ..... E. callistum (2)

40' Pedicels glabrous; flowers glabrous or hairy

41. Involucres, stems tomentose to (glabrous); leaf blades densely white- to yellow-brown-tomentose on both surfaces, or tomentose to glabrous adaxially; subshrub or mat — NCo, n&c CCo ..... E. latifolium

41' Involucres, stems glabrous, if tomentose then plant not in NCo, n&c CCo; leaf blades tomentose to densely so on both surfaces or tomentose to glabrous adaxially; plant generally erect ..... E. nudum (2)

42. Involucres, inflorescence branches tomentose to sparsely so; leaves generally basal

43. Fls white to rose or yellow; stem tomentose to hairy; leaf blades 2–4 cm; NW, CaR, n SN ..... var. oblongifolium

43' Fls white; stem densely tomentose (to tomentose); leaf blades 2–3.5 cm; s SNF ..... var. regirivum (2)

42' Involucres, inflorescence branches glabrous, if not then leaves sheathing

44. Lvs sheathing, margins generally strongly wavy

45. Sts tomentose

46. Fls hairy, 1.5–2 mm; involucres 3–4 mm; s SNF ..... var. regirivum (2)

46' Fls glabrous, 2–4 mm; involucres 3–6 mm; SnFrB

47. Lf blades (1)3–8 cm, (1)2–4 cm wide, sheathing 1–5(10) cm; c SnFrB ..... var. auriculatum (2)

47' Lf blades 1–3 cm, 1–1.5 cm wide, sheathing 5–20 cm; s SnFrB ..... var. decurrens

45' Sts glabrous

48. Lf blades densely tomentose abaxially, tomentose adaxially; involucres 5–10; s SNF ..... var. murinum

48' Lf blades tomentose abaxially, less so to glabrous adaxially; involucres 1–5; NW, CW

49. Involucres solitary; stem inflated; flowers white to yellow, glabrous; e SCoRI ..... var. indictum

49' Involucres 1–5 on same plant; stem not or ± inflated; flowers white to pink (or ± yellow), glabrous or hairy; NW, CW

50. Involucres (2)3–5 per cluster; stem not or ± inflated; flowers white to pink (or ± yellow), glabrous; NW, CW ..... var. auriculatum (2)

50' Involucres 1–3; stem not inflated; flowers white to pink, hairy; deltaic GV (Contra Costa Co.) ..... var. psychicola

44' Lvs basal, margins not or ± wavy

51. Involucres 1(2)

52. St inflated; flowers yellow (or white) — s SN, Teh, s CW, TR, s SNE, w DMoj ..... var. westonii (2)

52' St not inflated, if so then plant of PR; flowers white (or yellow)

53. Fls hairy (or glabrous); TR, PR ..... var. pauciflorum (2)

53' Fls glabrous; NW, CaR, SN, SnFrB, GB

54. Lf blades 1–2 cm; SNH, GB ..... var. deductum

54' Lf blades 1–5 cm; NW, CaR, SN, SnFrB ..... var. nudum (2)

51' Involucres 2–10

55. Fls hairy, generally yellow

56. St not inflated; leaf blades glabrous or woolly-tufted adaxially, margins flat; NW, CaR, SN, ScV, SnFrB ..... var. pubiflorum

56' St ± to distinctly inflated; leaf blades tomentose to sparsely so adaxially, margins wavy; s SNE, w DMoj ..... var. westonii (2)

55' Fls generally glabrous, white (or yellow)

57. Infl generally head-like or ± so; alpine, c&s SNH ..... var. scapigerum

57' Infl cyme-like, if head-like or ± so then not alpine

58. Involucres 5–7 mm, 1(2); TR, PR ..... var. pauciflorum (2)

58' Involucres 3–5 mm, 2–10; NW, CaR, SN, SnFrB

59. Infl cyme-like, branched 2+ times; involucres 2–5; NW, CaR, SN, SnFrB ..... var. nudum (2)

59' Infl head- or cyme-like, branched 1–2 times; involucres 5–10; n NCo ..... var. paralinum

34' Shrub

60. Lf blades narrowly linear to oblanceolate, <= 2 cm, generally strongly clustered; widespread ..... E. fasciculatum

61. Lvs thinly white-tomentose abaxially, glabrous adaxially; flowers, involucres generally glabrous; plant generally decumbent; CCo ..... var. fasciculatum

61' Lvs, flowers, involucres hairy, if ± glabrous then plant of deserts; plant erect to rounded; CA-FP, SNE, D

62. Lvs light yellow-green, glabrous or ± so adaxially; involucres, flowers glabrous or occasionally hairy; D ..... var. flavoviride

62' Lvs dark green or ± gray, generally hairy adaxially; involucres, flowers hairy; CA-FP, SNE, D

63. Infl open, mostly cyme-like; leaves densely white-tomentose abaxially, less so to green, sparsely tomentose adaxially, margins generally tightly rolled under; CW, SW, occasionally NW, D ..... var. foliolosum

63' Infl head- to umbel-like (or cyme-like); leaves gray-hairy on both surfaces or densely gray-tomentose abaxially, gray-hairy adaxially, margins not or occasionally rolled under; s SN, SCoRI, e SCo, TR, e PR, SNE, D ..... var. polifolium

60' Lf blades linear-oblong to round, if linear then > 2 cm and not clustered; CW, SW

64. Fls glabrous; CCo, SCo ..... E. parvifolium

64' Fls long-soft-hairy; CCo, SCo, ChI, occasionally established elsewhere

65. Lf blades linear to narrowly oblong, 2–4(5) cm, white-tomentose abaxially, gray-hairy to ± glabrous adaxially, margins rolled under; n ChI, occasionally established elsewhere ..... E. arborescens

65' Lf blades ovate or lanceolate to oblong- or narrow-ovate, 1.5–10 cm, white- or gray-tomentose generally on both surfaces, margins flat; CW, SW

66. Lf blades ovate, 1.5–3 cm, 1–2.5(3) cm wide; inflorescence generally head-like; s CCo, w SCo, n ChI (Santa Rosa Island), occasionally established CW, SW ..... E. cinereum

66' Lf blades oblong-ovate to ovate or lanceolate to narrowly oblong, 2–7(10) cm, 1–5 cm wide; inflorescence generally open, cyme-like; s ChI, occasionally established CW, SW ..... E. giganteum

67. Lf blades lance-oblong to lanceolate; San Clemente Island ..... var. formosum

67' Lf blades oblong-ovate to ovate; Santa Catalina, Santa Barbara islands

68. Lf blades 2.5–3.5(6) cm, 1.5–2(4) cm wide; inflorescence rarely open; Santa Barbara Island ..... var. compactum

68' Lf blades 3–7(10) cm, 2–5 cm wide; inflorescence open; Santa Catalina Island, occasionally CW, SW ..... var. giganteum

30' Involucres 1 per node

69. Involucres in raceme-like arrangements at tips of inflorescence branches

70. Involucres (4)6–7 mm; perennial herb, 6–12(18) dm — c&s CW, SW ..... E. elongatum var. elongatum

70' Involucres (0.8)1–6 mm; herb to shrub, (1)1.5–5 dm

71. Shrub — s MP, SNE ..... E. nummulare

71' Subshrub or herb

72. Plant basally ± woody, much-branched; leaf blades oblanceolate to elliptic, 0.1–1 cm wide, margins often rolled under

73. Involucres (4)5–6 mm; flowers (3)4–5 mm, cream to rose; n SCoRO ..... E. butterworthianum

73' Involucres 0.7–4 mm; flowers 1.5–4 mm, white to pink or rose; widespread ..... E. wrightii

74. Herb, loosely to compactly matted

75. Plant 0.1–0.3(0.6) dm; leaf blades 0.1–0.25 cm; involucres 0.8–1.7(2) mm; s SNH ..... var. olanchense (2)

75' Plant 0.5–2.5(3) dm; leaf blades 0.5–1(1.2) cm; involucres 1.5–4 mm; SN, CW, TR, e PR ..... var. subscaposum

74' Shrub or subshrub

76. Sts, inflorescence ± gray, tomentose to densely so — e PR, s DMoj, w DSon ..... var. nodosum

76' Sts, inflorescence ± white, ± red, or ± green, tomentose to woolly-tufted

77. Petiole bases forming distinct ring around stem; leaf blades 0.2–0.6(1) cm, 0.1–0.3(0.4) cm wide; SW ..... var. membranaceum

77' Petiole bases not forming ring around stem; leaf blades 0.5–3 cm, 0.2–1 cm wide; NW, s CaR, n SNF, DMoj

78. Lf blades 1.5–3 cm, 0.5–1 cm wide; involucres 2–4 mm; flowers 2.5–4 mm; NW, s CaR, n SNF ..... var. trachygonum

78' Lf blades 0.5–1.5 cm, 0.2–0.5(0.7) cm wide; involucres 2–2.5 mm; flowers 2.5–3.5 mm; DMoj ..... var. wrightii

72' Plant basally not woody, little-branched; leaf blades round to widely ovate, (0.3)1–3.5(4) cm wide, margins flat — W&I, DMtns

79. Lf blades ± round, (0.3)0.5–1.5 cm, petioles 1–2 cm; involucres bell-shaped ..... E. mensicola

79' Lf blades ovate to elliptic, oblong, or obovate, 1.5–4.5(5) cm, petioles 1–7 cm; involucres generally narrower than bell-shaped

80. Lf blades ovate to elliptic or obovate, 1.5–4 cm, 1–2.5 cm wide, petioles 1–5 cm; involucres 3–5 mm, 2–4 mm wide ..... E. panamintense

80' Lf blades elliptic to oblong, (2)2.5–4.5(5) cm, 1.5–3.5(4) cm wide, petioles (2)3–7 cm; involucres 3–4 mm, 3–4 mm wide ..... E. rupinum

69' Involucres in forks of branches

81. Herb

82. Lf blades ovate to elliptic or oblanceolate, (0.3)1–4 cm wide; CA (see 41' above) ..... E. nudum (2)

82' Lf blades round-ovate, 0.3–0.5(1) cm wide; n SNF ..... E. apricum

83. Sts erect to ± spreading ..... var. apricum

83' Sts prostrate ..... var. prostratum

81' Subshrub to shrub

84. Fls hairy

85. Fls green-yellow to yellow; involucres 4-toothed — s DSon ..... E. deserticola

85' Fls white; involucres 5–8-toothed

86. Involucres 3.5–5 mm; nw KR ..... E. pendulum

86' Involucres 2–3 mm; CaRH ..... E. spectabile

84' Fls glabrous

87. Infl panicle- or cyme-like, generally extensively branched, generally open, not terminal; involucres glabrous

88. Outer perianth lobes obovate; inflorescence branches ± gray, generally spreading, tiered; D ..... E. plumatella

88' Outer perianth lobes obovate to round; inflorescence branches green, ascending, not tiered; c&s SNH, Teh, s SCoR, TR, SNE, DMoj ..... E. heermannii

89. Sts, inflorescence branches round or angled, minutely scabrous or papillate-scabrous; plant generally densely branched; DMtns

90. Sts, inflorescence branches round, glabrous or minutely scabrous ..... var. argense

90' Sts, inflorescence branches sharply angled, deeply grooved, minutely scabrous ..... var. sulcatum

89' Sts, inflorescence branches round, not angled, generally smooth, glabrous or sparsely tomentose to woolly-tufted; plant sparsely branched; widespread

91. Involucres not in raceme-like arrangement, or only uppermost 2–3 so disposed; inflorescence branches glabrous — c&s SNH (e slope), SNE, n DMtns ..... var. humilius

91' Involucres in raceme-like arrangement, at least at tips of branches; inflorescence branches glabrous or sparsely tomentose to woolly-tufted

92. Infl sparsely tomentose to woolly-tufted; s CA ..... var. floccosum

92' Infl branches glabrous; s-c, sw CA

93. Lf blades 0.5–1.5 cm, glabrous to sparsely tomentose abaxially; inflorescence branches stout; s SNH (Kern Co.), Teh, se SCoRO (se San Luis Obispo Co.), n TR ..... var. heermannii

93' Lf blades 1.5–3(4) cm, generally tomentose to sparsely so abaxially (at least in early flower); inflorescence branches slender; c SCoRI ..... var. occidentale

87' Infl cyme-like, generally moderately branched, generally compact, terminal; involucres tomentose to glabrous

94. Plant matted subshrub, 0.5–1.5 dm; leaf blades 0.4–0.6 cm, margins rolled under; e DMtns ..... E. thornei

94' Plant erect to spreading subshrub or shrub, 0.2–15 dm; leaf blades (0.3)1–3.5 cm, margins flat, or if rolled under; widespread ..... E. microthecum

95. Fls yellow

96. Lf blades (0.2)0.3–0.6(0.8) cm wide; flowers (1.5)2–2.5(3) mm; involucres 2–2.5 mm; fruit 1.5–2 mm; SNH (e slope), GB ..... var. ambiguum

96' Lf blades 0.5–1.2 cm wide; flowers 2.5–3 mm; involucres 2.5–4 mm; fruit 2.5–3 mm; s MP ..... var. schoolcraftii

95' Fls white, cream, orange, pink, or red

97. Hairs ± white (see also Eriogonum microthecum var. alpinum) or stem, inflorescence rarely glabrous

98. Lf margins flat; inflorescence sparsely tomentose or glabrous; SN, GB ..... var. laxiflorum

98' Lf margins generally rolled under; inflorescence tomentose to densely so or rarely subglabrous; s SNE, DMoj ..... var. simpsonii

97' Hairs ± brown or ± red (generally white in Eriogonum microthecum var. alpinum) or stems, inflorescence generally glabrous

99. Shrub 3–6 dm

100. Sts, inflorescence tomentose to densely so; flowers 2–2.5(3) mm; fruit 2.5–3 mm; e SnBr ..... var. corymbosoides

100' Sts, inflorescence tomentose in youth, sparsely tomentose in age; flowers 1.5–2(2.5) mm; fruit 1.8–2 mm; s W&I, n DMtns ..... var. panamintense

99' Subshrub 0.2–1.5(2) dm

101. Lf blades elliptic or ovate, margins flat; flowers (1.5)2–3.5(4) mm

102. Lf blades 0.5–1 cm, (0.2)0.3–0.5(0.6) cm wide; involucres (2)2.5–3 mm; flowers (2.5)3–3.5(4) mm; e SnGb, w SnBr ..... var. johnstonii

102' Lf blades 0.3–0.7(0.8) cm, 0.1–0.4. cm wide; involucres 2.5–3.5 mm; flowers (1.5)2–3.5 mm; s W&I ..... var. lapidicola

101' Lf blades linear or linear-oblanceolate to narrowly elliptic, margins often rolled under; flowers 1.5–2.5 mm

103. Involucres (1.5)2–3 mm; flowers white to pink or rose; n&c SNH, n SNE ..... var. alpinum

103' Involucres 3–4 mm; flowers cream; SnBr ..... var. lacus-ursi

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].
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