Mitthyridium fasciculatum

Click the yellow dot to see these synapomorphies (click any yellow dot to see a clade's characters)

Cancellinae characters:
Cancellinae cell columns evolve from17 - 28

Gemmae characters: gemmae present.

Costa characters: abaxial epidermis evolves

Indel characters: indel 5 evolves.

Mitthyridium fasciculatum

Range Map

This clade is found throughout Malesia extending as far east as Samoa and as far west as Madagascar

Description

Lineage: Mitthyridium fasciculatum

Latin name: Mitthyridium fasciculatum (Hook. & Grev) Robinson
Synonyms:
Mitthyridium fasciculatum var. cardotii (Fleisch) A. Eddy
Mitthyridium fasciculatum var. rhizophyllum (Broth. & Park) Nowak
Syrrhopodon fasciculatus Hook. & Grev.
Thyridium fasciculatum (Hook. & Grev) Mitt.
<specimens examined 277, 283, 385, 369, 278, 292, 246, 804>

General features:
Plants robust, flat green to yellow-green in color, with densely foliate primary stems up to 20 cm long. Often pendant and on bark of trees, rarely on rocks. Leaves with suberect sheathing bases, and strongly flared at shoulders. Sporophytes frequent.

This group is found throughout Malesia extending as far east as Samoa and as far west as Madagascar. Generally the largest member of Mitthyridium and among the most abundant. The average elevation = 100 meters (N=200). Rarely found above 300m. Personal collections were from Fiji, Samoa, Australia, Sarawak (Borneo), and Peninsular Malaysia.

Diagnostic phylogenetic characters: [move the cursor over the image to view the morphological matrix]
Cancellinae cell columns are many, typically 25-30. The shape of the transition between cancellinae and lamina is typically flat. Margin serrations are present and usually large and extensive. The number of marginal border cell columns is on average 14. The marginal border tapers at approximately 1/3-1/4 the total leaf length. Leaf base rhizoids are very common. Leaves are commonly undulate. Papillae are common, large (obscuring laminal cell outlines) and many (3-4) per cell. Papillae are found on the dorsal side of the costa.
Diagnostic phylogenetic characters (synapomorphies and/or autapomorphies) were assessed by optimizing via ACCTRAN morphological characters to the 80 taxon total evidence phylogeny decribed in the Phylomonography&Mitthyridium section bulleted above.

Gemmae frequently present.
Gemmae are rough.
Gemmipars present.
Cancellinae – on average 30 cell columns on each side of the midrib.
Cancellinae composed on average 1/3 the total leaf length.
Hyaline border - on average, 8-12 marginal border cell columns.
Transition shape between lamina and cancellinae /\.
Leaf apex acute.
Adaxial epidermis present.
Abaxial epidermis weak or not present.
Dorsal stereid band present.
Ventral stereid band present.
Costa shape elliptical.
Large marginal serrrations present.
Margin tapers from 4 to 2 cells thick at 1/4 of the leaf length measured from the apex.
Margin more or less flat and unstratose.
Margin serrations cover the majority of the leaf (except at the cancellinae).
Leaf tip rhizoids rarely present.
Leaf base rhizoids frequent.
Leaf undulate.
Papillae present on the adaxial side of the costa.
Papillae cover laminal cell sufaces.
Papillae obscure the cell outlines in transverse sectional view.
Two or more papillae per laminal cell.
Autapomorphies (diagnostic characters): 4, 6, 19, 20, 21, 23