Moss Data Availablity Matrix

Complied and maintained by C. La Farge

The families are listed alphabetically within the classes following Vitt (1984), Vitt et al. (1998). Vitt's classification of the genera has been modified for a number of families: Pottiaceae Zander (1983), Rhacitheciaceae (Goffinet 1997), Fabroniaceae (Buck 1981).

This Matrix includes 5 levels of priority:

1. selected taxa to represent the major lineages of mosses (classes)
2. taxa to represent the orders or suborders, taxa easily sampled (17)
3. 30 major families represented
4. All major families (84) represented by 2 species
5. Monotypic families, all families represented

For the morphological data, preliminary columns have been outlined and some sample entries are shown (under the peristome development and blepharoplast columns) for how the entries should look. We would greatly appreciate any suggestions and additions for this part of the matrix.

**Note: The accession numbers are placed in the same order as the columnar order of genomic regions.

Acknowledgments to:

Bill Buck
Virginia Bryan
Ingrid Capesius
Russell Chapman
Cymon Cox
Efrain De Luna
Bernard Goffinet
Terry Hedderson
Jaakko Hyvvnen
Catherine La Farge
James Manhart
Brent Mishler
Angela Newton
Ron Pursell
Yin-Long Qui
Stephan Schaffer
Jon Shaw
Michael Stech
Dale Vitt
Allison Withey

Taxon

PRIORITY

DIVERS

specimen

Herb

Contact

Access

rRNA

cpDNA

cpDNA

cpDNA

nDNA

nDNA

PERIST

Morphology

BLEPHAROBLAST

SPERM

SPORE

SPORE

SPORE

SPORE

FAM/GEN

Voucher

Person

number

18S

rps4

trnL

rbcL

ITS

26S

DEV/ORN

WHOLE

SEM

TEM

DEV

GERM

Genbank

SPORLING

EMBL

SPHAGNOPSIDA

1

Sphagnaceae

1

Sphagnum cuspidatum

3

Capesius

X80213*

x

Sphagnum palustre

1

Qui 94018

Indiana

Chapman

x

Dukett & Carothers 1979

ANDREAOPSIDA

3

Andreaeaceae

1

Andreaea rothii

1

Andreaea rupestris

3

Hedderson 10439

RNG

Chapman

U18490

x

Andreaeobryaceae

1

Andeaeobryum macrospora

2

Takakiaceae

2

Takakia lepidozioides

1

Schofield

UBC

Chapman

U18526

x

POLYTRICHOPSIDA

3

Buxbaumiaceae

5

Buxbaumia aphylla

2

Diphyscium foliosum

2

Soderstrom SN 24/7/96

RNG

Cox/ Hedderson

x

x

x

Anderson & OToole 24708; Anderson 24714

DUKE

Shaw

Shaw et al. 1987

Polytrichaceae

21

Polytrichum commune

1

?

?

?

x

Hedderson 10458

RNG

Chapman

U18518

x

Polytrichum juniperinum

Paolillo & al. 1986a

Alophosia azorica

3

Rumsey SN

RNG

Cox/ Hedderson

x

Tetraphidaceae

2

Tetraphis pellucida

2

Hedderson 10454

RNG

Chapman

U18527

x

Anderson & Mishler 24,533; Anderson & O'Toole 24711

DUKE

Shaw

Shaw & Anderson 1988

BRYOPSIDA

84

Amblystegiaceae

22

Amblystegium serpens

Marcotte 8551

MICH

Shaw

Shaw & Rohrer 1984

Cox/Newton

x

x

?

DeLuna/Mishler

Calliergonella cuspidata

3

Stech

HEID

Capesius/ Stech

X94257

x

Calliergon giganteum

4

Archidiaceae

1

Archidium stellatum

5

?

Mishler

x

Archidium tennerimum

5

Brown & Carothers 1986

Aulacomniaceae

1

Aulacomnium androgynum

Cox 149

RNG

Cox/ Hedderson

AF023688; AF023811;AF023728

x

x

x

Aulacomnium palustre

4

Garriguees

MICH

Shaw

Shaw & Rohrer 1984

Aulacomnium heterostichum

5

Blomquist &Robertson 1941

Bartramiaceae

9

Bartramia stricta

3

Longton 4871

RNG

Cox/ Hedderson

AF023698;AF023799;AF023756

x

x

x

Bartramia crispata

4

Saito &Shimoze 1955

Philonotis fontana

2

Cox 117

RNG

Cox/ Hedderson

AF023694;AF023801;AF023758

x

x

x

Brachytheciaceae

35

Brachythecium rutabulum

3

Qiu 94071

UCB

Chapman

x

Stech

HEID

Capesius/ Stech

X94256

x

Hedderson 11763

RNG

Cox/ Hedderson

AF023713;AF023818;AF023772

x

x

x

Kindbergia praelongum

4

Hedderson 11765

RNG

Cox/ Hedderson

x

Eurhynchium pulchellum

4

Bryaceae

20

Brachymenium pulchrum

4

Perold

LSU

x

Swartz 2730

RNG

x

x

x

Bryum alpinum

4

Bryum bicolor

2

Shaw sn, 1986, Shaw sn 1986

DUKE

Shaw

Shaw et al. 1989

Pohlia nutans

3

Hedderson 10481 & 8869

RNG

Cox/ Hedderson

x

x

Rhodobryum roseum

3

Hedderson SN

RNG

Cox/ Hedderson

x

x

Bryobartramiaceae

1

Bryobartramia robinsii

5

Bryoxiphiaceae

1

Bryoxiphium norvegicum

5

Wallace SN 83

RNG

Cox/ Hedderson

x

Calomniaceae

1

Calomnion laetum

5

Vitt 29964

ALTA

Chapman

x

Calymperaceae

11

Syrrhopodon fimbriatulus

3

?

?

Mishler

x

Octoblepharum albidum

4

?

?

Mishler

x

Catoscopiaceae

1

Catoscopium nigritum

5

Longton 4592

RNG

Cox/ Hedderson

x

x

Chryphyaceae

8

Chryphaea patens

?

DeLuna

x

Chryphaea tasmanica

4

Acrochryphaea gardneri

4

Climaceaceae

1

Climacium dendroides

5

Cyrtopodaceae

3

Bescherellia elegantissima

4

Withey 732

DUKE

Withey/Mishler

x

Cryptopodium bartramioides

4

Glenny 4963

DUKE

Withey/Mishler

x

Cyrtopus setosus

4

Glenny 4827

DUKE

Withey/Mishler

x

Daltoniaceae

13

Acidontium megalocarpum

4

Richards R5830

MICH

Shaw

Shaw & Rohrer 1984

Daltonia tenuifolia

3

Dicnemonaceae

5

Dicnemon semicryptum

4

Streimann 51311

NY

La Farge

x

Eucamptodon muelleri

4

Streimann 55710

NY

La Farge

x

Dicranaceae

58

Campylopus introflexus

3

Hedderson 11768

RNG

Cox/ Hedderson

x

Dicranoloma billardii

3

La Farge-England 7515

ALTA

La Farge

x

Dicranum scoparium

2

?

?

Capesius

X89874*

x

?

?

Mishler

x

Qiu 94072

UCB

Chapman

x

Mishler 3757, 3759

DUKE

Shaw

Shaw et al. 1989

Leucoloma rutenbergii

3

La Farge-England 6939

ALTA

La Farge

x

Disceliaceae

1

Discelium nudum

5

Ditrichaceae

26

Ceratodon purpureus

4

Belland & Schofield 17530

DUKE

La Farge

x

Evans& Hooker 1913

Distichium inclinatum

4

Schofield 92239

DUKE

La Farge

x

Ditrichum flexicaule

4

Belland & Schofield 18296

DUKE

La Farge

x

Ditrichum pallidum

4

Hedderson 10350

Chapman

x

Anderson 24682, Shaw sn , 1987

DUKE

Shaw

Shaw et al. 1989

Echinodiaceae

1

Echinodium hispidum

5

Encalyptaceae

2

Bryobrittonia longipes

3

Vitt 18/6/1989

ALTA

Chapman

x

Vitt s.n.

ALTA

Cox/ Hedderson

AF023679;AF023778;AF023718

x

x

x

Encalypta ciliata

4

Mishler

x

Encalypta procera

2

Vitt 37966

ALTA

Chapman

x

Vitt 37966

ALTA

Goffinet

x

Entodontaceae

4

Erythrodontium squarrosum

4

Entodon beyrichii

4

?

DeLuna

x

Ephemeraceae

2

Ephemerum serratum

3

Schwartz 1994

Micromitrium tenerum

3

Ephemeropsaceae

1

Ephemeropsis tjibodensis

5

Erpodiaceae

4

Venturiella sp.

3

Aulacopilum hodgkinsoniae

4

Vitt 28261

ALTA

Goffinet

x

Eustichiaceae

1

Eustichia jamesonii

5

Fabroniaceae

3

Anacamptodon splachnoides

Anderson & Palmer 1982

Fabronia ciliaris

4

Leveriella perserrata

4

Fissidentaceae

1

Fissidens osmundoides

4

?

Mishler

x

Fissidens taxifolius

3

Stech

HEID

Capesius/ Stech

x

Cox 151

RNG

Cox/ Hedderson

x

Fissidens limbatus

Mueller 1973,1974

Fontinalaceae

4

Fontinalis antipyretica

4

Hedderson 11849

RNG

Cox/ Hedderson

AF023714;AF023817;AF023771

x

x

x

Dichelyma falcatum

4

Funariaceae

17

Funaria hygrometrica

1

?

Capesius

x

Schwartz 1994

Carother & Brown 1985

?

Mishler

x

Cox 148

RNG

Cox/ Hedderson

AF023776;AF023716

x

x

Shaw sn, 1986, Shaw sn, 1986

DUKE

Shaw

Shaw et al. 1989

Physcomitrium pyriforme

3

Hedderson 10352

RNG

Chapman

U18517

Schwartz 1994

Gigaspermaceae

4

Gigaspermum repens

4

Lorenziella imbricata

4

Grimmiaceae

12

Coscinodon cribosus

4

?

PMAE

Chapman

x

Grimmia torquata

3

Hedderson 11783

RNG

Cox/ Hedderson

x

Racomitrium fasciculare

2

?

Mishler

x

Hedderson 11804

RNG

Cox/ Hedderson

x

Schaffer 12

Schaffer (priv.)

Schaffer

x

Schistidium apocarpum

4

?

Mishler

x

Cox 154

RNG

Cox/ Hedderson

x

Scouleria aquatica

5

?

Mishler

x

Hedderson 5811

RNG

Cox/ Hedderson

AF023684;AF023780;AF023723

x

x

x

Newberry 5077

Newberry (priv.)

Schaffer

x

Hedwigiaceae

6

Hedwigia ciliata

2

?

Capesius

x

Goffinet 3324

ALTA

Goffinet

x

Rhacocarpus purpurascens

4

Hedderson 11790

RNG

Cox/ Hedderson

N/A;AF023815;AF023724

x

x

x

?

Capesius

X89869*

x

Heliocophyllaceae

1

Heliocophyllum torquatum

5

Hookeriaceae

27

Hookeria lucens

2

Cox 118

RNG

Cox/ Hedderson

x

Brymelaparkeriana

3

?

DeLuna

x

Hydropogonaceae

2

Hydropogon fontinaloides

4

Hydropogonella gymnostoma

4

Hylocomiaceae

12

Hylocomium splendens

4

Stech

HEID

Capesius/ Stech

x

Pleurozium schreberi

4

Hedderson 11767

RNG

Cox/ Hedderson

x

?

Mishler

x

Hypnaceae

40

Hypnum cupressiforme

3

Stech

HEID

Capesius/ Stech

X94258

x

Hedderson 11764

RNG

Cox/ Hedderson

x

Hypnum mamillatum

Duckett & Renzaglia 1986

Ptilium crista-castrensis

4

Hypnodendraceae

22

Braithwaitea sulcata

4

Streimann 38403

RNG

Cox/ Hedderson

AF023820;AF023745

x

x

Hypnodendron comosum

3

Hypopterygiaceae

5

Hypopterygium tahitense

3

Withey 520

DUKE

Withey/Mishler

x

Lopidium struthiopteris

4

Lembophyllaceae

3

Camptochaete arbuscula

4

Lembophyllum divulsum

4

Leptodontaceae

Leptodon smithii

?

DeLuna/Mishler

x

Leptostomataceae

1

Leptostomum macrocarpum

5

Fletcher SN

RNG

Cox/ Hedderson

AF023696;AF023790;AF023744

x

x

x

Leptostomum inclinans

5

Lepyrodontaceae

1

Lepyrodon tomentosus

5

?

DeLuna

x

Leskeaceae

27

Leskea angustata

DeLuna/Mishler

x

Leskea radicosa

3

Lindbergia brachyptera

4

Leucobryaceae

4

Brothera leana

4

Leucobryum albidum

4

Qui 94075

UCB

Chapman

x

Schaffer 12

Schaffer (priv.)

Schaffer

x

Leucodontaceae

12

Leucodon julacea

3

?

Mishler

Leucodon assimilis

4

Hedderson 11768

RNG

Cox/ Hedderson

x

Antitrichia curtipendula

4

Meesiaceae

3

Amblyodon dealbatus

4

Hedderson 3975a

RNG

Cox/ Hedderson

AF023806;AF023731

x

x

Meesia uliginosa

4

Hedderson 6481

RNG

Cox/ Hedderson

AF023693;AF023803;AF023735

x

x

x

Meteoraceae

19

Meteorium teres

4

?

De Luna

x

Papillaria deppei

4

?

De Luna

x

Microtheciellaceae

1

Microtheciella kerrii

5

Mitteniaceae

1

Mittenia plumula

5

Shaw 1985

Mniaceae

9

Koponen 1969

Cinclidium stygium

4

Hedderson 10477

RNG

Cox/ Hedderson

AF023706;AF023791;AF023763

x

x

x

Mnium hornum

2

?

Capesius

X80985*

x

Cox 115

RNG

Cox/ Hedderson

AF023796;AF023767

x

x

Plagiomnium affine

3

Hedderson 10461

RNG

Cox/ Hedderson

AF023711;AF023797;AF023762

x

x

x

Myrinaceae

6

Schwetschkeopsis denticulata

4

Myrinia pulvinata

4

Helicodontium capillare

4

?

DeLuna/Mishler

x

Nanobryaceae

1

Nanobryum drummondii

5

Neckeraceae

22

Metaneckera menziesii

4

Hedderson 5738

TAH

Cox/ Hedderson

x

Thamnobryum alopecurum

4

Cox 147

RNG

N/A;AF023834; AF023769

x

x

x

Oedipodiaceae

1

Oedipodium griffithianum

5

Duckett SN

RNG

Cox/ Hedderson

x

Orthotrichaceae

21

/Lewinsky 1989 (17spp)

Macrocoma tenuisssp. tenuis

4

Breedlove 69342

ALTA

Goffinet

x

Orthotrichum obtusifolium

2

Vitt 33870

ALTA

Goffinet

x

Schotheimia rugifolia

3

Zygodon reinwardtii

4

Goffinet 636

ALTA

Goffinet

x

/Lewinsky 1990 (10 spp)

Zygodon pungens

3

La Farge-England 8097* cf pungens

ALTA

Goffinet

x

/Lewinsky 1990 (10 spp)

Phyllodrepaniaceae

2

Phyllodrepanium ?

5

Phyllogoniaceae

4

Catagonium politum

4

Phyllogonium vicosum

4

Plagiotheciaceae

10

Plagiothecium undulatum

4

Stech

HEID

Capesius/ Stech

x

Stereophyllum radicosulum

4

Pleurophascaceae

1

Pleurophascum grandiglobum

5

Pleuroziopsaceae

1

Pleuroziopsis n.n.

5

Pottiaceae

76

Zander, 1993

Stegonia latifolia

3

Timmiella anomala

2

Hedderson 9434

RNG

Cox/ Hedderson

x

Tortula ruralis

3

Cox 120

RNG

Cox/ Hedderson

AF023682;AF023831;AF023722

x

x

x

Priodontaceae

3

Prionodon densus

?

DeLuna

x

Prionodon fusco-lutescens

4

Neolindbergia robusta

4

Pseudoditrichaceae

1

Pseudoditrichium buchananii

5

Pterobryaceae

29

Garovaglia elegans

?

DeLuna/Mishler

x

Garovaglia subelegans

3

Baker 740

RNG

Cox/ Hedderson

x

x

Hildebrandtiella pachyclada

5

Pterobryon densum

?

DeLuna

x

Ptychomitriaceae

4

Glyphomitrium cylindraceum

4

Ptychomitrium gardneri

4

Hasse

ALTA

Chapman

x

Ireland 7038

PMAE

Goffinet

x

Goffinet 580 *

Cox/ Hedderson

AF023689;AF023779;AF023719

x

x

x

Ptychomniaceae

7

Ptychomnion aciculare

?

DeLuna/Mishler

x

Regamatodontaceae

1

Regamatodon filiformis

5

Rhabdoweisiaceae

4

Rhabdoweisia crenulata

4

Amphidium lapponicum

4

Rhachitheciaceae

7

Uleastrum palmicola

Vitt 21161

ALTA

Goffinet

x

Rhacithecium papillosum

4

Hypnodontopsis mexicana

4

Racopilaceae

2

Racopilum tomentosum

3

?

DeLuna/mishler

x

Schnepf et al. 1978

Racopilum convolutaceum

4

Glenny 4941

DUKE

Withey/Mishler

x

Powellia involutifolia

3

Rhizogoniaceae

10

Goniobryum subbasilare

4

Streimann 38105

RNG

Cox/ Hedderson

AF023824;AF023753

x

x

Pyrrhobryum spiniforme

3

Pocs & al. 88110/BJ

RNG

Cox/ Hedderson

AF023832;AF023751

x

x

Rhytidiaceae

1

Rhytidium rugosum

5

?

DeLuna

x

Rutenbergiaceae

2

Neorutenbergia armata

4

Rutenbergia madagassa

4

?

DeLuna

Schistostegaceae

1

Schistostega pennata

5

Seligeriaceae

7

Blinda acuta

3

Hedderson 11828

RNG

Cox/ Hedderson

AF023681;AF023781;AF023721

x

x

x

Brachydontium tichodes

4

Anon SN 82

RNG

Cox/ Hedderson

x

Seligeria pusilla

3

Hattaway 957

UCB

Schaffer

x

Sematophyllaceae

50

Brotherella recurvens

4

Renzaglia 865

Manhart

L13475

x

Glossadelphus laevifolius

4

Sorapillaceae

1

Sorapilla sprucei

5

?

DeLuna/mishler

x

Splachnaceae

8

Koponen 1982

Tayloria lingulata

2

Tetraplodon mnioides

3

Goward 95-1470

ALTA

Goffinet

x

Schwartz 1994

Splachnum amplulaceum

4

Schwartz 1994

Spiridentaceae

2

Franciella spiridentoides

4

Withey 756

DUKE

Withey/Mishler

x

Spiridens reinwardtii

4

Eddy 6232

BM

Cox/ Hedderson

AF023828;AF023748

x

x

Thuidiaceae

19

Abietinella abietina

2

Goffinet 4106

ALTA

Goffinet

x

Thuidium tamariscinum

3

Hedderson 11766

RNG

Cox/ Hedderson

AF023819;AF023770

x

x

Timmiaceae

1

Timmia austriaca

2

Hedderson 5684

RNG

Cox/ Hedderson

x

Timmia sibirica

4

Hedderson 6341

RNG

Cox/ Hedderson

AF023678;AF023775;AF023715

x

x

x

Trachypodaceae

6

Trachypodopsis oranans

4

?

DeLuna/Mishler

x

Trachypus bicolor

4

Viridivelleraceae

1

Viridivellus pulchellum

5

Wardiaceae

1

Wardia hygrometrica

5

Hedderson 11709

RNG

Cox/ Hedderson

AF023683;AF023782;AF023720

x

x

x