GREEN PLANT PHYLOGENY RESEARCH COORDINATION GROUP
TABLE 4 - Last update 11/12/96
DATA AVAILABILITY MATRIX FOR FERNS, SPHENOPSIDS AND PSILOTACEAE
This table originally compiled by P. R. Crane, J. A. Doyle, P. Gensel, W. Hahn, P. Kenrick, R. Olmstead, K. Pryer, A. Smith, D. Soltis, E. A. Zimmer
at the 1995 Berkeley meeting of the GPPRCG and modified based on broader input at the 1996 Seatttle meeting

q possibly very incomplete; + information; - no information; ?AP questionable association of parts (for fossils)
Taxon - fossils indicated # 
Priority
clade represented and extant diversity
extant
diversity
sporophyte vegetative morphology
sporophyte vegetative anatomy
sporangia
/seeds
spores
gametophyte
sperm 
ultrastruct.
rbcL
atpB
18S
other genes
 comments 
Sphenopsids 
ca. 15 sp.
Equisetum hyemale 
2
Sphenopsid
ca. 15 sp.
+
+
+
+
+
?
-
-
U18500
26S: Zimmer (partial)
represents subg. Hippochaete
Equisetum arvense 
1
Sphenopsid
ca. 15 sp.
+
+
+
+
+
?
L11053
Wolf
-
16S: U24593
represents subg. Equisetum
#Archaeocalamites 
1
Sphenopsid
-
+
+
+ ?AP
+
-
-
-
-
-
-
-
#Calamites 
2
Sphenopsid
-
+
+
+
+
-
-
-
-
-
-
-
#Sphenophyllum 
1
Probable Sphenopsid
-
+
+
+
+
-
-
-
-
-
-
-
Psilotaceae 
ca. 12 sp.
Tmesipteris tannensis 
2
Psilotaceae
ca. 12 sp.
+
+
+
+
+
?
-
Wolf
U18103
-
-
Psilotum nudum 
1
Psilotaceae
ca. 12 sp.
+
+
+
+
+
?
L11059
Wolf
X81963
16S: U24590; 26S: Zimmer (partial)
-
Eusporangiate ferns - Marattiales 
ca. 200 sp.
Angiopteris evecta 
1
Marattiaceae
ca. 200 sp.
+
+
+
+
+
?
L11052
Wolf
-
16S: U24580
-
Marattia salicina 
2
Marattiaceae
ca. 200 sp.
+
+
+
+
+
?
-
-
-
-
-
Christensenia aesculifolia 
2
Marattiaceae
ca. 200 sp.
+
+
+
+
+
?
-
-
-
-
-
Danaea nodosa 
2
Marattiaceae
ca. 200 sp.
+
+
+
+
+
?
-
-
-
-
-
Eusporangiate ferns - Ophiogloss. 
ca. 80 sp.
Botrychium strictum 
2
Ophioglossaceae
ca. 80 sp.
+
+
+
+
+
?
B14881
-
-
-
represents subg. Japanobotrychium
Botrychium lunaria 
2
Ophioglossaceae
ca. 80 sp.
+
+
+
+
+
?
L40966
Wolf
U18623
-
represents subg. Botrychium
Botrychium multifidum 
1
Ophioglossaceae
ca. 80 sp.
+
+
+
+
+
?
-
-
-
-
represents subg. Sceptridium
Botrychium viginianum 
3
Ophioglossaceae
ca. 80 sp.
+
+
+
+
+
?
-
-
-
-
represents subg. Osmundopteris
Ophioglossum engelmannii 
2
Ophioglossaceae
ca. 80 sp.
+
+
+
+
+
?
L11058
-
-
16S: U24589
-
Basal leptosporangiate ferns 
ca. 1000 sp.
Osmunda cinnamomea 
1
Osmundaceae
ca. 18 sp.
+
+
+
+
+
?
D14882
Wolf
U18516
16S: U24594
-
Trichomanes davallioides 
1
Hymenophyllaceae
ca. 600 sp.
+
+
+
+
+
?
U05948
Wolf
U18629
26S: Wolf (partial)
sometimes put in Vandenboschia or Cephalomanes
Cephalomanes thysanostomum 
2
Hymenophyllaceae
ca. 600 sp.
+
+
+
+
+
?
U05608
-
-
-
-
Dipteris conjugata 
1
Dipteridaceae
ca. 7 sp.
+
+
+
+
+
?
U05620
-
-
-
-
Cheiropleuria bicuspis 
2
Cheiropleuriaceae
ca. 1 sp.
+
+
+
+
+
?
U05607
-
-
-
-
Stromatopteris moniliformis 
2
Gleicheniaceae
ca. 125 sp.
+
+
+
+
+
?
U05653
-
-
-
-
Dicranopteris linearis 
1
Gleicheniaceae
ca. 125 sp.
+
+
+
+
+
?
U18626
-
U18625
-
-
Diplopterygium glaucum 
2
Gleicheniaceae
ca. 125 sp.
+
+
+
+
+
?
U05624
-
-
-
-
Matonia pectinata 
1
Matoniaceae
ca. 4 sp.
+
+
+
+
+
?
U05634
-
-
-
-
Actinostachys digitata 
2
Schizaeaceae
ca. 170 sp.
+
+
+
+
+
?
U05650
-
-
-
-
Anemia phyllitidis 
1
Schizaeaceae
ca. 170 sp.
+
+
+
+
+
?
-
-
-
-
sometimes put in Anemiaceae
Lygodium japonicum 
2
Schizaeaceae
ca. 170 sp.
+
+
+
+
+
?
U05632
-
-
16S: U24588
sometimes put in Lygodiaceae
Tree ferns 
ca. 700 sp.
Dicksonia antarctica 
1
Dicksoniaceae
ca. 40 sp.
+
+
+
+
+
?
U05919
Wolf
U18624
-
-
Calochlaena dubia 
2
Dicksoniaceae
ca. 40 sp.
+
+
+
+
+
?
U05912
-
-
-
-
Plagiogyria japonica 
2
Plagiogyriaceae
ca. 25 sp.
+
+
+
+
+
?
U05643
-
-
-
-
Sphaeropteris lepifera 
2
Cyatheaceae
ca. 650 sp.
+
+
+
+
+
?
U05616
-
-
-
-
Metaxya rostrata 
2
Metaxyaceae
ca. 1 sp.
+
+
+
+
+
?
U05635
-
-
-
-
Water ferns 
ca. 80 sp.
Marsilea quadrifolia 
1
Marsileaceae
ca. 65 sp.
+
+
+
+
+
?
U05633
-
-
-
-
#Hydropteris pinnata 
2
fossil intermediate 
-
+
-
+
+
-
-
-
-
-
-
-
Salvinia cucullata 
2
Salviniales.
ca. 16 sp
+
+
+
+
+
?
U05649
-
-
-
-
Salvinia natans 
2
Salviniales
ca. 16 sp.
+
+
+
+
+
?
-
-
X90413
-
-
Azolla caroliniana 
2
Salviniales
ca. 16 sp.
+
+
+
+
+
?
U24185
-
-
-
-
Pteridoid ferns 
ca. 1000 sp.
Ceratopteris richardii 
1
Parkeriaceae
ca. 4 sp.
+
+
+
+
+
?
-
-
-
-
model system
Vittaria flexuosa 
2
Vittariaceae
ca. 110 sp.
+
+
+
+
+
?
U05656
-
-
-
-
Coniogramme japonica 
2
Pteridaceae
ca. 845 sp.
+
+
+
+
+
?
U05611
-
-
-
-
Adiantum raddianum 
2
Pteridaceae
ca. 845 sp.
+
+
+
+
+
?
U05906
Wolf
U18621
-
-
Adiantum capillus-veneris 
1
Pteridaceae
ca. 845 sp.
-
-
-
-
+
?
U14880
Wolf
-
-
-
Pteris fauriei 
2
Pteridaceae
ca. 845 sp.
+
+
+
+
+
?
U05647
-
-
-
-
Acrostichum aureum 
2
Pteridaceae
ca. 845 sp.
+
+
+
+
+
?
U05601
-
-
-
-
Taenitis blechnoides 
2
Pteridaceae
ca. 845 sp.
+
+
+
+
+
?
U05654
-
-
-
-
Platyzoma microphyllum 
2
Pteridaceae
ca. 845 sp.
+
+
+
+
+
?
U05644
-
-
-
-
Dennstaedtioid ferns 
ca. 400 sp.
Lindsaea odorata 
2
Dennstaedtiaceae
ca. 400 sp.
+
+
+
+
+
?
U05630
-
-
-
sometimes put in Lindsaeaceae
Pteridium aquilinum 
1
Dennstaedtiaceae
ca. 400 sp.
-
-
-
-
+
?
U05646
Wolf
-
-
also rbcL: U05939
Microlepia strigosa 
2
Dennstaedtiaceae
ca. 400 sp.
+
+
+
+
+
?
U05931
-
-
-
-
Dennstaedtia punctilobula 
2
Dennstaedtiaceae
ca. 400 sp.
+
+
+
+
+
?
U05918
Wolf
-
-
-
Monachosorum henryi 
2
Dennstaedtiaceae
ca. 400 sp.
+
+
+
+
+
?
U05932
-
-
-
-
Histiopteris incisa 
2
Dennstaedtiaceae
ca. 400 sp.
+
+
+
+
+
?
U05627
-
U18630
-
-
Blotiella pubescens 
2
Dennstaedtiaceae
ca. 400 sp.
+
+
+
+
+
?
U05911
-
-
-
-
Lonchitis hirsuta 
2
Dennstaedtiaceae
ca. 400 sp.
+
+
+
+
+
?
U05929
Wolf
U18632
-
-
Polypodioid ferns 
ca. 1150 sp.
Polypodium australe 
1
Polypodiaceae
ca. 665 sp.
+
+
+
+
+
?
U21140
-
-
-
-
Loxogramme grammitoides 
2
Polypodiaceae
ca. 665 sp.
+
+
+
+
+
?
U05631
-
-
-
-
Micropolypodium okuboi 
2
Grammitidaceae
ca. 450 sp.
+
+
+
+
+
?
U05658
-
-
-
-
All other derived ferns 
ca. 5000 sp.
Asplenium filipes 
2
Aspleniaceae
ca. 720 sp.
+
+
+
+
+
?
U05907
Wolf
-
-
-
Thelypteris dentata 
1
Thelypteridaceae
ca. 915 sp.
+
+
+
+
+
?
-
-
-
-
-
Thelypteris beddomei 
2
Thelypteridaceae
ca. 915 sp.
+
+
+
+
+
?
U05655
-
-
-
-
Blechnum occidentale 
2
Blechnaceae
ca. 240 sp.
+
+
+
+
+
?
U05909
-
-
26S: Wolf (partial)
-
Blechnum brasiliense 
2
Blechnaceae
ca. 240 sp.
+
+
+
+
+
?
U05910
Wolf
U18622
-
-
Onoclea sensibilis 
2
Dryopteridaceae
ca. 1045 sp.
+
+
+
+
+
?
U05936
-
-
-
sometimes put in Athyriaceae
Rumohra adiantiformis 
2
Dryopteridaceae
ca. 1045 sp.
+
+
+
+
+
?
U05942
-
-
-
-
Elaphoglossum hybridum 
2
Dryopteridaceae
ca. 1045 sp.
+
+
+
+
+
?
U05924
-
-
-
sometimes put in Lomariopsidaceae
Nephrolepis cordifolia 
2
Dryopteridaceae
ca. 1045 sp.
+
+
+
+
+
?
U05933
-
-
-
sometimes put in Nephrolepidaceae
Davallia mariesii 
2
Dryopteridaceae
ca. 1045 sp.
+
+
+
+
+
?
U05617
-
-
-
sometimes put in Davalliaceae