#NEXUS [MacClade 3.01 registered to Equator, UCMP] BEGIN DATA; DIMENSIONS NTAX=27 NCHAR=113; [!This is the final revised morphological matrix for land plants (LP-MORPH), corrected 12/93, per comments of co-authors. **used in the Ann .Mo. Bot. 81: 451-483 (Mishler et al. 1994) green plant synthesis paper. Characters are re-numbered as they are used in our text, but you can see Karen Renzaglia's old numbers (from the sperm data set, Garbary et al. 1993, Pl. Syst. & Evol., which see for more details on characters), or reference to other literature (M&C = Mishler and Churchill, 1985, Cladistics 1:305-328) as footnotes to the characters.] FORMAT MISSING=? GAP=- SYMBOLS= " 0 1 2 3 4 5 6"; OPTIONS MSTAXA=UNCERTAIN ; CHARLABELS [1] apical_cell_in_antheridia [2] div._pat._in_young_antheridia [3] endogenous_antheridia [4] antheridial_stalk [5] operculum_cells [6] sperm_in_pollen_tube [7] number_of_sperm_per_male_struc. [8] nascent_spermatids [9] diagonal_spindle_in_final_mitot [10] replication_of_the_centrioles [11] time_of_origin_of_centrioles [12] basal_bodies_and_flagella [13] bicentrioles [14] basal_body_position [15] proximal_extension_A [16] proximal_extension_B [17] stellate_transition [18] connect._fibers_betw._basal_bod [19] basal_body_structure [20] BB_staggering_assoc._w._MT_grow [21] regression_of_LS [22] 'LS/AM_elongation_longit.' [23] spline_aperture [24] sline_aperture_location [25] position_of_developing_MLS [26] plaque_stratified_betw._blephar [27] 'spline/LS_orientation' [28] posterior_notch_to_LS [29] LS_position [30] stray_spline_MT [31] accessory_band_of_MTs [32] maturational_elongation_of_AM [33] spline_shank [34] osmiophilic_crest [35] anterior_osmiophilic_crest [36] changes_in_BBs_at_mat. [37] matrix_around_BBs [38] posterior_of_the_stellate_patt. [39] flagellar_scales [40] late_blepharoplast_w._transient [41] direction_of_flagellar_emergenc [42] nuclear_shape_at_maturity [43] nuclear_posterior_shape [44] median_constriction [45] splin_attached_to_nucleus [46] spline_growth_assoc._w._nucl._s [47] direction_of_nuclear_compaction [48] condensed_chromatin_strands [49] diverticulum_during_shaping [50] number_of_gyres_of_nucleus [51] dense_body_in_ant._mitochondrio [52] mit._assoc._w._plastids_in_sper [53] mit._assoc._w._plastids_in_sper [54] specialized_ant._mito. [55] specialized_posterior_mito. [56] additional_mito._in_ant._of_cel [57] origin_of_AM [58] osmophilic_material_underneath [59] change_from_cristae_sacs_to_baf [60] monoplastidic_sperm [61] plastid_determines_division_pol [62] starch_grains_in_single_plastid [63] sperm_plastid_contacting_nucleu [64] fibrillenscheide [65] cytoplasmic_loss [66] embryo [67] cuticle [68] lunularic_acid [69] elaters [70] oil_bodies [71] 'D-Methionine' [72] stomates [73] vert_div_of_zygote [74] xylem [75] phloem [76] perine_layer_on_spores [77] 'aerial_sporophyte_axis_(persist' [78] columella [79] Multicellular_rhizoids [80] leaves_on_gam. [81] independent_sporophyte [82] branched_sporophyte [83] tracheids [84] lignin [85] 'long_exserted_seta_(of_liv._typ' [86] oil_body_cells [87] spore_mother_cells_lobed [88] 'capsule_2-4_valved' [89] 'caps_wall_cells_w/_trans_thick' [90] thick_caps._wall [91] elongate_antheridia [92] aerial_calyptra [93] paraphyses [94] costate_leaves [95] peristome [96] operculum [97] cylindical_sprogen_layer [98] transfer_cells_on_gam_side [99] transfer_cells_on_spor.__side [100] seed [101] megaphyll [102] microphyll [103] 'lateral,_broad_sporangia' [104] exarch_maturation_of_xylem [105] cp_DNA_inversion [106] sporangia_borne_on_leaves [107] trichomes [108] flavonoids [109] retention_of_zygote [110] sheathed_hairs [111] polyphenolics_induced_by_sex [112] parenchyma [113] beaked_muscilage_papillae ; STATELABELS 1 absent present, 2 'four-celled' 'two-celled', 3 absent present, 4 absent present, 5 absent present, 6 absent present, 7 1000+ '100-1000' '16-24' 2, 8 paired not_paired, 9 absent present, 10 present absent, 11 always_present sperm._mother_cells sperm._moth._cell_progenitor earlier, 12 two more_than_two, 13 present absent, 14 right_angles 'side-by-side' 'staggered_ant-post' staggered_continous, 15 absent long short, 16 'ventral_-_dorsal' ventral, 17 present absent, 18 present absent fine_filaments_w._centrin, 19 monomorphic dimorphic, 20 absent present, 21 absent complete partial, 22 parallel perpendicular, 23 absent present, 24 left_of_center right_of_center, 25 adjacent_to_BBs beneath_BBs, 26 absent present, 27 90_deg. 45_deg., 28 absent present, 29 under_all_BBs under_ABB_only under_some_BBs, 30 absent present develops_late, 31 absent present, 32 absent posterior, 33 wide less_than_4_tubules, 34 absent present, 35 absent present, 36 absent dense_material_at_tip BB_cartwheel_w._plug BB_triplets_impreg._w._matrixma, 37 homogenous mottled, 38 extracell._or_partly entirely_intracell., 39 present absent, 40 yes no, 41 toward_side toward_rear toward_anterior, 42 ovoid elongate, 43 not_expanded expanded, 44 absent present, 45 yes detached_at_maturity never_attached, 46 absent present, 47 outer_shell anterior_to_posterior at_equal_rates_along_nucl. general_increase_in_density, 48 'spaghetti-like' perpendicular_to_spline 'spiral-central_strand' general_compaction spikes irregular_plates solid_mass_from_ant._tip, 49 absent present, 50 not_coiled '0.5-3' greater_than_3, 51 absent present, 52 absent present, 53 absent present, 54 present absent, 55 present absent, 56 absent row_of_mito._behind_AM numerous_unspecialized, 57 fusion elongation, 58 absent present, 59 absent present, 60 present absent, 61 'present_-_at_poles' 'present_-_asymmetrical' absent, 62 more_than_one one, 63 absent present, 64 absent present, 65 absent partial 'complete_(or_tiny_remn.)', 66 absent present, 67 absent present, 68 absent present, 69 absent present, 70 absent present, 71 not_recognize recognize, 72 absent present, 73 absent present, 74 absent present, 75 absent present, 76 absent present, 77 absent present, 78 absent present, 79 absent present, 80 absent 'present_(of_moss_type)' 'present_(of_junger._type)', 81 absent present, 82 absent present, 83 absent present, 84 absent present, 85 absent present, 86 absent present, 87 absent present, 88 absent present, 89 absent present, 90 'absent_(unistrat.)' 'present_(2-10_strat.)', 91 absent present, 92 absent present, 93 absent present, 94 absent present, 95 absent present, 96 absent present, 97 absent present, 98 absent present, 99 absent present, 100 absent present, 101 absent present, 102 absent present, 103 absent present, 104 absent present, 105 absent present, 106 absent present, 107 absent present, 108 absent present, 109 absent present, 110 absent present, 111 absent present, 112 absent present, 113 absent present, ; MATRIX CHARA ?????0??0030?00?020???0??0???00?0000000?11000000010??1?1?011201020000000000000000000000000?0000000000000?00100000 NITELL ?????0??0030?00?020???0?000??00?0000000?11000000010001???0?1201020000000000000000000000000?0000000000000?00100000 COLEOC_PUL ?????0???000?00?0?0?000?000??0000000000?00?00???000000?1??0000?010000000000000000000000000?0000000000000?00011100 COLEOC_ORB ?????0??1000?00?0?0?000?000??0000000000?00?00???000000?1??0000?010000000000000000000000000?0000001000000?00011110 PHAEOCEROS 0011000001101111100?200?101000010003001?1101001101111000011011102110001110000100000000000000000001000000?00010110 NOTOTHYLAS 0011000001101111100?200?101000010003001?1101001101111000011011102110001110000100000000000000000001000000?00010110 MARCHANTIA 000100000110121001112011101100000003001?1100001011001000001010112111110000000000000001000000000001100000000110110 SPHAEROCARPOS 000100000110121001112011101100001003001?1100001011001000001010??211111000000000000000{01}000000000001100000?00110110 PELLIA 010100000110121001112011101000000003001?1100101011001000001010112111110000000000000010111100000000000000?00110110 BLASIA 0?0100000110121001112011101100000003001?1100001011001000001010112111110000000000000010111100000001100000?00110110 JUNGERMANNIA 010100000110121001112011101000000003001?1100101011001000001010112111110000000002000010111100000000100000?00110110 HAPLOMITRIUM 010100000110121?01112010101000?00?03001?11000015?10010000010?1??2111110000000000000010110000000001100000?00110110 TREUBIA 01010000011012100111?010101000?00?03?01?11?0001???0??0000??0?0??211111000000000000001011110000000??00000?00110110 SPHAGNUM 110100011110121011111011101012010003001?1100012201011000001001002110001100011111000000000001000100000000?00010110 ANDREAEA 1101100111101210?1112011101010010003001?1100012201011000001000002110001001011111000000000011110000100000?00010111 POLYTRICHUM 110110011110121001112011101011011003001?1100012201011000001000002110001101111111000000000011111110100000?00010110 HYPNUM 110110011110121001112011101011010003001?1100012211011000001000002110001101111111000000000011111111100000000110110 TAKAKIA 11011001111012100111?01110101??10?03?01?1100?????10?10000?1000??2110001001??11?10000000000110?0000100000?00110111 LYCOPODIELLA 0?00001?1110140?020?010?1010000?0?00101?10?000??00001010?000?0001110001101111000111100000000000001100111000110110 LYCOPODIUM 0?00001011101221020?010?101000010000101?10000?3310000010?0100010{12}110001101111000111100000000000001100111000110110 SELAGINELLA 0??0?01??110?2210?0?200?101000010000101?110000?401001000?010?000211000110111100011110000000000000??00111000110110 EQUISETUM 0?0000111111030?0100010?1110201001100011111000330100001110012?00111000110111100011110000000000000??00?00110110110 PTERIDIUM 0?000021112103210?00010?111020100112?011111000330200001110012?001110001111111000111100000000000001101000111110110 MARSILEA 0??0?0211121030?0100010?1110200000020011110000160200001010012?00111000111111100011110000000000000??01000?11110110 OSMUNDA 0?000021112103210?00010?1110201001110011111000330100001110012?001110001111111000111100000000000001101000111110110 GINKGO ?????13?11?1030?0100010?1?1020000110011020?02???00000112?0012?0001100011011?100011110000000000000??11000111110110 ZAMIA ?????13?11?1030?0100010?1?1020000110011020?02???00000112?0012?0001100011011?100011110000000000000??11000111110110 ; END; BEGIN ASSUMPTIONS; OPTIONS DEFTYPE=unord PolyTcount=MINSTEPS ; TYPESET * order_two = unord: 1-6 8-49 51-113, ord: 7 50; WTSET * CURRENT = 1: 1-113; EXSET sperm_only = 1-65; EXSET general_morph = 66-113; END; begin paup; constraint moss+trach = (1,2,3,4,5,6,7,8,9,10,11,12,13,(14,15,16,17,18,19,20,21,22,23,24,25,26,27)); constraint horn+moss+trach = (1,2,3,4,7,8,9,10,11,12,13,(5,6,14,15,16,17,18,19,20,21,22,23,24,25,26,27)); constraint bryos = (1,2,3,4,(19,20,21,22,23,24,25,26,27,(5,6,7,8,9,10,11,12,13,14,15,16,17,18))); end; BEGIN NOTES; TEXT CHARACTER=1 TEXT= 'KAREN''S_#2'; TEXT CHARACTER=2 TEXT= 'KAREN''S_#3'; TEXT CHARACTER=3 TEXT= 'KAREN''S_#5'; TEXT CHARACTER=4 TEXT= 'KAREN''S_#6'; TEXT CHARACTER=5 TEXT= 'KAREN''S_#7'; TEXT CHARACTER=6 TEXT= 'KAREN''S_#8'; TEXT CHARACTER=7 TEXT= 'KAREN''S_#10__!ORD'; TEXT CHARACTER=8 TEXT= 'KAREN''S_#12'; TEXT CHARACTER=9 TEXT= 'KAREN''S_#14'; TEXT TAXON=20 TEXT= 'Lycopodium_obscurum_added_from_Karen''s_notes_sent_3/1/93_--_note_that_the_old_Lycop_is_now_Lycopodiella_!'; TEXT CHARACTER=10 TEXT= 'KAREN''S_#15'; TEXT CHARACTER=11 TEXT= 'KAREN''S_#16'; TEXT CHARACTER=12 TEXT= 'KAREN''S_#17 '; TEXT CHARACTER=13 TEXT= 'KAREN''S_#18'; TEXT CHARACTER=14 TEXT= 'KAREN''S_#19 '; TEXT CHARACTER=15 TEXT= 'KAREN''S_#20'; TEXT CHARACTER=16 TEXT= 'KAREN''S_#21'; TEXT CHARACTER=17 TEXT= 'KAREN''S_#22'; TEXT CHARACTER=18 TEXT= 'KAREN''S_#24'; TEXT CHARACTER=19 TEXT= 'KAREN''S_#25'; TEXT CHARACTER=20 TEXT= 'KAREN''S_#26'; TEXT CHARACTER=21 TEXT= 'KAREN''S_#27'; TEXT CHARACTER=22 TEXT= 'KAREN''S_#28'; TEXT CHARACTER=23 TEXT= 'KAREN''S_#30'; TEXT CHARACTER=24 TEXT= 'KAREN''S_#31'; TEXT CHARACTER=25 TEXT= 'KAREN''S_#32'; TEXT CHARACTER=26 TEXT= 'KAREN''S_#34'; TEXT CHARACTER=27 TEXT= 'KAREN''S_#35'; TEXT CHARACTER=28 TEXT= 'KAREN''S_#37'; TEXT CHARACTER=29 TEXT= 'KAREN''S_#38'; TEXT CHARACTER=30 TEXT= 'KAREN''S_#39'; TEXT CHARACTER=31 TEXT= 'KAREN''S_#40'; TEXT CHARACTER=32 TEXT= 'KAREN''S_#44'; TEXT CHARACTER=33 TEXT= 'KAREN''S_#47'; TEXT CHARACTER=34 TEXT= 'KAREN''S_#49'; TEXT CHARACTER=35 TEXT= 'KAREN''S_#50'; TEXT CHARACTER=36 TEXT= 'KAREN''S_#51'; TEXT CHARACTER=37 TEXT= 'KAREN''S_#52'; TEXT CHARACTER=38 TEXT= 'KAREN''S_#53'; TEXT CHARACTER=39 TEXT= 'KAREN''S_#54'; TEXT CHARACTER=40 TEXT= 'KAREN''S_#55'; TEXT CHARACTER=41 TEXT= 'KAREN''S_#57'; TEXT CHARACTER=42 TEXT= 'KAREN''S_#58'; TEXT CHARACTER=43 TEXT= 'KAREN''S_#59'; TEXT CHARACTER=44 TEXT= 'KAREN''S_#61'; TEXT CHARACTER=45 TEXT= 'KAREN''S_#63'; TEXT CHARACTER=46 TEXT= 'KAREN''S_#64'; TEXT CHARACTER=47 TEXT= 'KAREN''S_#65'; TEXT CHARACTER=48 TEXT= 'KAREN''S_#66'; TEXT CHARACTER=49 TEXT= 'KAREN''S_#67 '; TEXT CHARACTER=50 TEXT= 'KAREN''S_#69__ORD!'; TEXT CHARACTER=51 TEXT= 'KAREN''S_#70'; TEXT CHARACTER=52 TEXT= 'KAREN''S_#71'; TEXT CHARACTER=53 TEXT= 'KAREN''S_#72'; TEXT CHARACTER=54 TEXT= 'KAREN''S_#74'; TEXT CHARACTER=55 TEXT= 'new_character_added_by_Karen_3/1/93'; TEXT CHARACTER=56 TEXT= 'KAREN''S_#75'; TEXT CHARACTER=57 TEXT= 'KAREN''S_#76'; TEXT CHARACTER=58 TEXT= 'KAREN''S_#77'; TEXT CHARACTER=59 TEXT= 'KAREN''S_#79'; TEXT CHARACTER=60 TEXT= 'KAREN''S_#80'; TEXT CHARACTER=61 TEXT= 'KAREN''S_#81'; TEXT CHARACTER=62 TEXT= 'KAREN''S_#83'; TEXT CHARACTER=63 TEXT= 'KAREN''S_#84'; TEXT CHARACTER=64 TEXT= 'KAREN''S_#86'; TEXT CHARACTER=65 TEXT= 'KAREN''S_#89'; TEXT CHARACTER=77 TEXT= 'of_moss/tracheophyte_type'; TEXT CHARACTER=80 TEXT= unord.; TEXT CHARACTER=85 TEXT= 'of_the_liverwort_type_(non-homologous_to_those_of_the_moss/tracheophyte_lineage)__--__M&C_char._#9'; TEXT CHARACTER=88 TEXT= 'on_a_regular_basis_(Andreaea_considered_non-homologous)_--_M&C_char._#14'; TEXT CHARACTER=108 TEXT= see_Markham_in_Zinsmeister_&_Mues; TEXT CHARACTER=90 TEXT= 'caps._wall_2-10_cells_thick_--__M&C_char._#19_--_thick_caps._walls_in_horn.,_mosses,_and_tracheo._considered_non-nomologous'; TEXT CHARACTER=89 TEXT= M&C_char._#18; TEXT CHARACTER=86 TEXT= M&C_char._#10; TEXT CHARACTER=87 TEXT= M&C_char._#13; TEXT TAXON=18 CHARACTER=80 TEXT= considered_homologous_with_mosses; TEXT TAXON=15 CHARACTER=94 TEXT= assuming_loss_in_some_spp.; TEXT TAXON=17 CHARACTER=94 TEXT= assuming_loss_in_some_spp.; TEXT TAXON=15 CHARACTER=74 TEXT= possible_homology_in_Andreobryum_?; TEXT CHARACTER=82 TEXT= with_multiple_sporangia; TEXT CHARACTER=109 TEXT= through_sporogenesis; TEXT TAXON=15 CHARACTER=72 TEXT= fide_Murray_1988; TEXT TAXON=15 CHARACTER=93 TEXT= fide_Murray_1988; TEXT TAXON=15 CHARACTER=113 TEXT= 'in_Andreaobryum_only_(fide_Murray_1988),_Andreaea_has_normal_moss-like_muscilage_papillae'; TEXT CHARACTER=113 TEXT= homology_accepted_from__Murray_1988; TEXT CHARACTER=106 TEXT= Bremer_char._#_64; TEXT CHARACTER=107 TEXT= Bremer_#_67; TEXT CHARACTER=105 TEXT= from_Raubeson_and_Jansen_1992; TEXT CHARACTER=103 TEXT= Crane; TEXT CHARACTER=104 TEXT= Crane; TEXT CHARACTER=102 TEXT= Crane; TEXT TAXON=4 CHARACTER=9 TEXT= not_sure_of_homology_with_mosses_+_tracheophytes; TEXT CHARACTER=74 TEXT= 'N.B.,_homology_between_mosses_and_tracheophytes_not_agreed_to_by_all_co-authors'; TEXT CHARACTER=75 TEXT= 'N.B.,_homology_between_mosses_and_tracheophytes_not_agreed_to_by_all_co-authors'; TEXT TAXON=8 CHARACTER=86 TEXT= 'polymorphic_in_order;__Riella_has_differentiated_cells'; TEXT TAXON=18 CHARACTER=94 TEXT= 'state_difficult_to_determine_--_maybe_all_costa!'; TEXT TAXON=22 CHARACTER=102 TEXT= not_sure_if_homologous_with_lycophytes; END; BEGIN MACCLADE; v 3.0 -1322664995 1100&/0 0 0 END;