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California Moss eFlora: General Key

· Home · List of Genera · Key to Keys · Accepted Names · Synonyms · For Beginners · Subdivisions of CA · Jepson eFlora for CA Vascular Plants


Wed Jan 23 07:45:11 2013



General Key.

A. Gametophyte essentially absent; plant visible as a large sporophyte (8 mm or more high) with a papillose seta and an asymmetric, capsule on an erect seta; restricted to rotten logs or highly organic soil .....Buxbaumia: B. piperi, B. viridis, B. aphylla
A. Not as above .....B

B. Leaves ecostate (plants considered to be ecostate usually have a short double costa that extends only a few cells above the leaf insertion; only a few cannot be shown to have a costa), or doubly costate with neither branch extending much above mid-leaf .....C
B. Leaves unicostate (plants considered unicostate generally have that costa extending to mid-leaf or beyond but sometimes that costa ends slightly below mid-leaf; some unicostate mosses have accessory short costae on each side of the main costa; some unicostate mosses may have that costa with short forks near its distal end) .....AH

C. Leaves with hyaline awns; leaves deeply concave .....D
C. Leaves without hyaline awns; leaves various, mostly not concave .....F

D. Plant erect and unbranched; central portion of leaf with photosynthetic filaments; capsules arranged on long and straight setae .....Aloina: A. bifrons
D. Plant prostrate and extensively branched central portion of leaf without photosynthetic lamellae; setae long or short .....E

E. Median cells strongly papillose; leaves longer than 1 mm; capsules mostly immersed on short lateral branches .....Hedwigia: Braunia secunda not known from CA, Hedwigia ciliata not known from CA, H. stellata, H. detonsa, Pseudobraunia californica
E. Median cells smooth; leaves very small, seldom more than 0.5 mm long; capsules exserted .....Lescuraea: Iwatsukiella leucotricha not known from CA

F. Median and upper leaf cells papillose or prorate .....G
F. Median and upper leaf cells smooth or, at most, with a few of the cells near the apex sparingly prorate .....K

G. Reddish to blackish erect and essentially unbranched plant growing in cushions on exposed siliceous rocks .....Andreaea: A. alpestris, A. rupestris
G. Plant extensively branched and mostly prostrate to decumbent or forming pinnate axes which may be partially ascending .....H

H. Paraphyllia present and obvious, especially on main stem axis .....I
H. Paraphyllia absent or leaf-like; branching not as above .....J

I. Plant regularly many times pinnately branched with the branches arranged in a single plane; leaves of branches markedly differentiated from those of the main stem .....Hylocomium: Hylocomium splendens
I. Plant sparingly and irregularly branched; branch and stem leaves very similar .....Lescuraea: Pterigynandrum filiforme

J. Costa double with both branches reaching nearly to mid-leaf; leaves large, mostly more than 4 mm long .....Hylocomium: Rhytidiadelphus triquetrus
J. Costa absent or short; leaves smaller .....Heterocladium: H. macounii, H. dimorphum, Myurella julacea, Pseudoleskeella serpentinensis, P. tectorum

K. Plant complanate-foliate .....L
K. Plant not at all complanate .....S

L. Leaves strongly undulate, serrate near apex with the distal teeth on some of the leaves curled back and pointing toward the leaf base .....Neckera: N. douglasii
L. Leaves entire to serrulate or, if somewhat serrate, the teeth not recurved and the leaves not undulate .....M

M. Median leaf cells mostly over 50 µm broad; leaf apices obtuse to broadly acute; nematogons present at leaf apices .....Hookeria: H. acutifolia not known from CA, H. lucens
M. Median leaf cells more narrow or the leaf without nematogons in the apical region .....N

N. Interior basal cells strongly porose; at least some leaves of the stem tips with serrate apices .....Hypnum: Herzogiella seligeri, H. striatella
N. Interior basal cells not porose; leaves entire to serrulate .....O

O. Outer cortical cells of stem thick-walled and narrow-lumened, thus resembling adjacent inner cortical cells .....P
O. Outer cortical cells of stem mostly larger and thinner-walled than adjacent inner cells, not resembling inner cortical cells .....Q

P. Stem and branch leaves radically different with the stem leaves long acuminate but the branch leaves bluntly acute to almost obtuse; median cells mostly less than 5:1 .....Heterocladium: H. procurrens not known from CA
P. Plant essentially isophyllous; median laminal cells mostly more than 8:1 .....Plagiothecium: Dacryophyllum falcifolium, Isopterygium tenerum, Pseudotaxiphyllum elegans

Q. Leaves at least somewhat concave with a short, usually forked costa or with a strong double costa; leaf apex blunt; plant aquatic or semiaquatic .....Hygrohypnum: H. ochraceum
Q. Leaves not at all concave; costa, if present, not forked above the base; leaf apex usually acute; plant of mesic habitats .....R

R. Leaves strongly decurrent to auriculate-decurrent .....Plagiothecium: all Plagiothecium, Buckiella undulata
R. Leaves not decurrent; tropical plant escaped as a lawn weed in coastal cities of California .....Hypnum: Vesicularia vesicularis

S. Leaf apex blunt, obtuse or rounded, never with an apiculus or an awn .....T
S. Leaf apex acute to acuminate or subulate .....Y

T. Plant erect and unbranched, with deeply concave leaves formed by infolded leaf margins, with photosynthetic lamellae arranged along the leaf center .....Aloina: A. aloides not known from CA, A. rigida
T. Plant prostrate and branched, without photosynthetic lamellae on the leaves .....U

U. Leaves 3-ranked and usually so strongly carinate (except in Fontinalis hypnoides) as to have the two lateral halves of the leaves pressed against one another; plant aquatic .....Fontinalis: all Fontinalis
U. Leaves not 3-ranked, never strongly carinate .....V

V. Alar cells so inflated as to appear swollen and overlapping, very hyaline, these cells making a group ascending 4-10 cells up the margin and a similar distance toward the middle; stem tips with leaves so closely appressed as to appear fusiform .....Hypnum: Calliergonella cuspidata
V. Alar cells less inflated and fewer; stem and branch apices not at all fusiform .....W

W. Leaves less than 0.5 mm long, coarsely dentate, closely arranged on julaceous stems .....Heterocladium: Myurella julacea
W. Leaves larger, entire to serrulate, not julaceous .....X

X. Plant of mesic to very wet but never submerged sites, mostly pinnately branched from obvious reddish stems; alar cells quadrate and mostly somewhat thick-walled .....Hylocomium: Pleurozium schreberi not known from CA
X. Plant aquatic or in constantly very wet sites, seldom pinnately branched, never with red stems; alar cells various .....Hygrohypnum: all Hygrohypnum

Y. Plant very small with larger leaves less than 1 mm long .....Z
Y. Plant and leaves larger .....AD

Z. Plant unbranched with a cleistocarpous, sessile capsule arising from a rosette of strongly serrate to fimbriate leaves; protonema usually forming a conspicuous greenish felt .....Acaulon: Ephemerum serratum
Z. Plant with stems more than l mm long; branched and usually prostrate; sporophyte, when present, stegocarpous .....AA

AA. Leaves dentate to fimbriate with their terminal cells abruptly several times as long as the adjacent subterminal ones .....Fabronia: F. ciliaris; F. pusilla
AA. Leaves with entire to serrulate or serrate margins; apical cells of leaves not so differentiated .....AB

AB. Plant julaceous .....Lescuraea: Pseudoleskeella serpentinensis, Tripterocladium leucocladulum
AB. Leaves spreading to loosely erect .....AC

AC. Leaves somewhat falcate-secund; costa forked distally below or near mid-leaf; cuticle often minutely verruculose .....Lescuraea: Leptopterigynandrum austro-alpinum; Pseudoleskeella tectorum
AC. Leaves symmetric with costa shorter and not-forked; cuticle smooth .....Amblystegium: Platydictya jungermannioides

AD. Plant whitish-green, densely fasciculate-branched; leaf cells of two forms with large, empty transparent ones alternating with small, green ones (these green cells may be so narrow as to be interpreted as the lateral walls of the transparent cells); plant restricted to very wet sites .....Sphagnum: all Sphagnum
AD. Not as above .....AE

AE. Leaves 3-6 mm long, broadly ovate, often plicate; costae two, with both extending to leaf middle or beyond .....Hylocomium: Rhytidiadelphus loreus, R. squarrosus, R. subpinnatus not known from CA, Rhytidiopsis robusta not known from CA
AE. Leaves smaller and usually ovate-lanceolate or more narrow; costae shorter .....AF

AF. Plant plumose and ascending from an inconspicuous prostrate base; alar cells very numerous, quadrate and pachydermous, extending more than 10 cells up the margin .....Antitrichia: Alsia californica, Pterogonium gracile
AF. Plant prostrate to ascending without strong differentiation of a prostrate main axis and an erect secondary axis; alar cells fewer and poorly differentiated .....AG

AG. Leaves squarrose but not falcate; abruptly contracted below or near the middle into a long and channeled acumen .....Campylium: C. hispidulum, C. stellatum
AG. Leaves not squarrose, often falcate-secund, with apices various but without a channeled acumen .....Hypnum: all Hypnum, Breidleria pratensis not known from CA, Homomallium mexicanum, Isopterygiopsis pulchella, Ptilium crista-castrensis not known from CA, Sematophyllum adnatum, Orthothecium chryseum not known from CA

AH. Basal and sometimes all cells thick-walled with the lateral walls sinuose .....Racomitrium: all Racomitrium
AH. Cells with lateral walls not sinuose .....AI

AI. Cells papillose, cuticular papillose, strongly mammillose or evenly prorate at least in the distal one-half of the limb .....AJ
AI. Cells smooth, or with a few scattered prorae in the distal portion of the leaf .....CE

AJ. Leaves with hyaline to yellowish awns .....AK
AJ. Leaves not awned or with the awns concolorous with the adjacent apex or only slightly bleached near the distal end .....AQ

AK. Plants prostrate and much branched; costa ending before the apex and the awn composed mostly of lamina cells .....Claopodium: C. bolanderi, C. crispifolium
AK. Plants erect and usually little branched; or, if branched, the awn formed from the excurrent costa .....AL

AL. Costa with adaxial photosynthetic lamellae or filaments; margins plane to incurved .....CU
AL. Costa without such outgrowths .....AM

AM. Plants with multicellular gemmae obvious in leaf axils or inserted on leaf lamina; leaf margins plane or nearly so; plants usually less than 5 mm high and mostly epiphytic .....Syntrichia: S. laevipila, S. pagorum, S. papillosa
AM. Plants without such multicellular gemmae; leaf margins and plant habit various .....AN

AN. Leaf margins plane to erect, or slightly recurved near middle; awn mostly smooth; calyptra large and campanulate enveloping all the capsule .....Encalypta: E. brevicollis not known from CA, E. brevipes not known from CA, E. ciliata, E. procera, E. rhaptocarpa, E. vulgaris
AN. Leaf margin recurved at least near base or, if with plane margins, with a serrulate awn; calyptra cucullate and sheathing only distal 1/2 of capsule .....AO

AO. Basal cells not enlarged and thin-walled; alar cells quadrate; adaxial surface of costa covered by rectangular cells with somewhat sinuose lateral walls .....Grimmia: G. elatior, G. lesherae, Schistidium strictum not known from CA
AO. Basal cells mostly large, rectangular and thin-walled without alar differentiation; adaxial surface of costa with quadrate cells similar to those of lamina .....AP

AP. Costa in cross-section with a single crescentic abaxial stereid band and mostly without a differentiated dorsal epidermis .....Syntrichia: S. bartramii, S. caninervis, S. laevipila, S. norvegica, S. obtusissima, S. papillosissima, S. princeps, S. ruralis
AP. Costa in cross-section with an elliptical abaxial stereid band and sometimes with a small adaxial band; epidermis of abaxial face of costa well-differentiated .....Tortula: T. brevipes, T. euryphylla, T. guepinii, T. muralis, Pseudocrossidium crinitum.

AQ. Leaf margins in distal portion of leaf serrulate to dentate .....AR
AQ. Leaf margins entire throughout, or with toothing only near the base, or within a few cells of the apex .....BK

AR. Plants branched .....AS
AR. Plants unbranched or branched only by innovations, sometimes seemingly irregularly branched .....AW

AS. Plants plumosely branched with the branches somewhat elevated above the substratum; leaves not catenulate when dry .....AT
AS. Plants prostrate; leaves catenulate when dry .....AV

AT. Alar cells thick-walled and very deeply green in an explanate pocket; most of the cells in the distal portion of the leaf prorate-spinose; plant mostly pendent from tree limbs in the northwest of the state .....Isothecium: I. spiculiferum
AT. Alar cells not so differentiated; habitat various .....AU

AU. Stems and branches with leaves julaceous; median cells prorate; alar cells quadrate, extending in a large group more than 10 cells up the leaf margin and toward the costa .....Antitrichia: Dendroalsia abietina, Pterogonium gracile
AU. Plants not julaceous; median cells unipapillose or pluripapillose; alar cells not uniformly quadrate .....Hylocomium: Helodium blandowii, Rhytidium rugosum not known from CA, Thuidium recognitum not known from CA

AV. Plants without foliose paraphyllia on stems and branches; cells of leaf margin almost without papillae; median cells centrally unipapillose .....Claopodium: C. whippleanum, Meteorium nigrescens not known from CA
AV. Plants mostly with such paraphyllia; cells of leaf margin with similar papillosity to those of interior; median cells mostly with papillae near cell ends .....Lescuraea: L. incurvata, L. pallida, L. patens, L. radicosa, L. saviana, L. stenophylla

AW. Leaf base differentiated with that base completely wrapped around stem, hyaline to reddish, strongly differentiated in cell shape and color from the limb .....AX
AW. Leaf base sometimes partially sheathing but never completely encircling the stem .....AY

AX. Leaves strongly mammillose with the limb rigidly appressed when dry .....Timmia: T. austriaca, T. bavarica, T. megapolitana not known from CA
AX. Leaves strongly papillose or prorate-papillose with the limb often somewhat crispate .....Anacolia: Bartramia ithyphylla

AY. Marginal cells differentiated as a limbidium, either elongate or epapillose and somewhat larger than adjacent isodiametric laminal cells .....Tortula: T. subulata, Hennediella heimii, H. stanfordensis
AY. Marginal cells not differentiated in the manner described above .....AZ

AZ. Alar cells very much inflated, usually reddish in older leaves; leaves falcate-secund .....Dicranum: D. fuscescens, D. sulcatum
AZ. Alar cells neither inflated nor strongly colored .....BA

BA. Median cells of leaves pluripapillose; cells across the entire leaf base rectangular, thin-walled, smooth and pellucid .....Barbula: Barbula ehrenbergii, Bryoerythrophyllum recurvirostrum
BA. Median cells of leaves unipapillose, prorate or with cuticular papillosity; basal cells various .....BB

BB. Leaves 10:1 or more, mostly subulate with costa filling most of the apex and percurrent to excurrent .....BC
BB. Leaves broader with costa of various lengths .....BG

BC. Median cells with cuticular papillose ridges running continuously from near the base to the apex; leaves crispate when dry .....BD
BC. Median cells without such cuticular ridges; leaves crispate or not .....BE

BD. Leaf margins remotely serrate to serrulate with very small teeth right angular in outline; leaves spirally inserted, not obviously ranked .....Amphidium: A. californicum
BD. Leaf margins evenly and coarsely dentate with pointed teeth; leaves in 3 straight ranks on the stem .....Anacolia: Plagiopus oederianus not known from CA

BE. Plant red-brown to almost black, growing on siliceous rock at high elevations, often below snow beds; capsules without an operculum, dehiscent by four longitudinal lines .....Andreaea: A. nivalis
BE. Leaf limb symmetrical to falcate; plant green to brownish green; habitat various; capsules dehiscent by operculum .....BF

BF. Leaf margins recurved; older portion on stem with dense red-brown, papillose rhizoids inserted in a halo around the macronematal apparatus .....Anacolia: A. baueri, A. laevisphaera, A. menziesii, Bartramia pomiformis, B. stricta, Plagiopus oederianus not known from CA
BF. Leaf margins plane to incurved; rhizoids smooth, not densely inserted and inserted around the macronematal apparatus .....Ditrichum: Distichium capillaceum, Distichium inclinatum, Ditrichum heteromallum, Ditrichum schimperi, Trichodon cylindricus

BG. Leaves bistratose with cells high mammillose only on adaxial surface; plant of low to mid-elevation, mostly disturbed soils .....Timmiella: T. anomala, T. crassinervis
BG. Leaves unistratose with cells papillose, mammillose or prorate .....BH

BH. Leaves in five distinct ranks with that ranking emphasized by the strong keeling of each leaf; plant mostly among other mosses at high elevations .....Philonotis: Conostomum tetragonum
BH. Leaves not ranked, not strongly keeled; habitat various .....BI

BI. At least some of the cells of the distal leaf margin prorate, mostly with all of the distal cells prorate; leaves lanceolate, 2.5-3.5:1; acute to acuminate at apices .....Philonotis: P. americana, P. caespitosa, P. calcarea, P. capillaris, P. fontana, P. marchica, P. muhlenbergii, P. tomentella, P. uncinata not known from CA, P. yezoana
BI. Median leaf cells mammillose to centrally unipapillose; leaves mostly ovate to elliptic, bluntly acute to obtuse at apices .....BJ

BJ. Leaf apices acute to acuminate; lower portion of plant densely covered with red rhizoids; median cells centrally unipapillose .....Aulacomnium: A. androgynum, A. palustre
BJ. Leaf apices bluntly acute to rounded; lower portion of plant not densely covered with rhizoids; median laminal cells mammillose or papillose on cellular mammillosities .....Dichodontium: Dichodontium flavescens, D. olympicum, D. pellucidum, Cynodontium strumulosum not known from CA

BK. Leaf margins plane to ascending or incurved, not recurved or with inconspicuous recurvature near the middle .....BL
BK. Leaf margins recurved, at least near the base .....BS

BL. Plant prostrate and branched; paraphyllia numerous, foliose .....Lescuraea: all Lescuraea, Leskea polycarpa
BL. Plant erect and sparsely or not at all branched; paraphyllia absent .....BM

BM. Papillae primarily of cuticular origin and thus not conforming positionally to cell outlines when viewed on leaf whole-mount .....Amphidium: A. lapponicum, Dicranoweisia contermina
BM. True papillae present .....BN

BN. Leaves ending in a uniseriate awn that may be bleached but is not truly hyaline; plants almost strictly epiphytic in California .....Ulota: Ulota megalospora, Zygodon rupestris
BN. Leaves variously acute or acuminate to obtuse at apex, never uniseriate at apex; ecology various .....BO

BO. Plant with elliptic-oblong leaves broadly rounded toward the apex; dry leaves not or little distorted, not truly crispate .....BP
BO. Plant with lanceolate leaves acute to acuminate at apex; dry leaves crispate or not .....BQ

BP. Median leaf cells pluripapillose; basal cells rectangular and inflated with transverse walls thin but with lateral walls somewhat thick and pigmented yellow to orange or reddish .....Encalypta: Encalypta affinis not known from CA, E. intermedia, E. procera, E. vulgaris
BP. Median leaf cells pluripapillose or unipapillose; basal cells rectangular and thick-walled, not pigmented but with some sinuosity on lateral walls .....Orthotrichum: O. bolanderi, O. obtusifolium

BQ. Basal cells of leaf very much inflated and rectangular, extending upward along margin significantly farther than in the juxtacostal region; the margin between the laminal and basal cells thus describing a ā€œVā€-shape .....Tortella: T. alpicola, T. fragilis, T. tortuosa
BQ. Basal marginal cells not extending distally with those basal cells not describing a "V" .....BR

BR. Margin strongly inrolled or infolded when dry; cells of limb rather uniform in size and shape .....Weissia: all Weissia, Oxystegus tenuirostris, Plaubelia sprengelii not known from CA, Trichostomum brachydontium, T. crispulum, T. sweetii
BR. Margin plane to erect; cells of strongly variable size and shape throughout the leaf limb .....Gymnostomum: Anoectangium aestivum not known from CA, Eucladium verticillatum, Gymnostomum aeruginosum not known from CA, G. calcareum, Molendoa sendtneriana, Zygodon rupestris

BS. Leaves with an obvious limbidium .....BT
BS. Leaves without a limbidium .....BV

BT. Limbidium formed of abruptly elongate cells .....Tortula: T. subulata
BT. Limbidium formed of cells larger, more pachydermous, or less papillose cells .....BU

BU. Plant large with leaves mostly more than 3 mm long; limbidium formed of one or several rows of larger and thicker, often reddened cells; plant usually growing in calcareous seeps .....Scouleria: Crumia latifolia
BU. Plant smaller with limbidium formed of one or several rows of epapillose and more pellucid cells; plant of temporarily moist soil .....Syntrichia: S. bolanderi

BV. Plants less than 3 mm high with the leaves closely inserted; costa percurrent to excurrent .....BW
BV. Plants mostly larger, not bulbiform .....BX

BW. Median cells with obvious C-shaped papillae; plant strongly bulbiform with an essentially sessile capsule .....Acaulon: Phascum cuspidatum
BW. Median cells with numerous papillae that appear round in surface view; plant bulbiform but with capsule emergent to exserted .....Microbryum: all Microbryum

BX. Papillae cuticular and therefore seen as low ridges that frequently extend across boundaries between adjacent cells; leaves when dry spirally coiled into several turns from apex to base .....Amphidium: A. mougeotii; Grimmia anomala
BX. Leaves with true papillae; mostly with dry leaves not spirally coiled .....BY

BY. Adaxial surface cells of costa rectangular with sinuose lateral walls; immediate suprabasal cells near costa strongly porose; some leaves in clone often with a hyaline awn .....Grimmia: G. lesherae
BY. Adaxial surface cells of costa various in shape but never with sinuose lateral walls; suprabasal cells near costa various; leaves mostly not with hyaline awns .....BZ

BZ. Costa homogeneous; papillae few, mostly high and sharp; capsules short elliptic, often sulcate; calyptra campanulate, usually hairy and plicate .....Orthotrichum: O. affine, O. alpestre, O. anomalum not known from CA, O. consimile, O. cupulatum, O. diaphanum, O. euryphyllum, O. flowersii, O. hallii, O. laevigatum, O. lyellii, O. macounii, O. pallens, O. papillosum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pylaisii, O. rivulare, O. rupestre, O. shevockii, O. speciosum, O. spjutii, O. striatum, O. tenellum, O. texanum, O. underwoodii, Ulota megalospora, U. obtusiuscula, U. phyllantha
BZ. Costa with at least a dorsal stereid and sometimes also with a ventral one; papillae low and numerous, obscuring the lumen; capsules longer cylindric, not sulcate; calyptra mostly cucullate .....CA

CA. Leaves elliptic to broadly ovate; costa cross-section without an adaxial stereid band and with abaxial stereid band lunate, without either an abaxial or an adaxial epidermis .....CB
CA. Leaves narrowly ovate to lanceolate; costa often with two stereid bands; abaxial stereid band elliptic .....CC

CB. Leaf margins revolute in more than one complete spiral; hydrated plants with a distinct pin-wheel appearance .....Didymodon: Pseudocrossidium obtusulum
CB. Leaf margins plane to incurved or recurved; hydrated plants otherwise .....Syntrichia: S. latifolia

CC. Central strand absent in stem cross-section; juxtacostal cells of median leaf slightly elongate, mostly with small trigones; peristome absent with operculum usually systylious .....Gymnostomum: Hymenostylium recurvirostrum
CC. Central strand usually present in stem cross-section; juxtacostal cells of median leaf similar to other median laminal cells; peristome present, mostly long and spirally twisted .....CD

CD. Cell lumina tending to be quadrate with the walls equally thickened throughout; papillae low and not obscuring the cell outlines, usually four or more per cell; axillary hairs with a basal brown cell; leaf apex mostly acute, often narrowly so; costa with at least a strong dorsal stereid .....Didymodon: Bryoerythrophyllum columbianum, B. recurvirostrum; Didymodon australasiae, D. brachyphyllus, D. eckeliae, Didymodon fallax, D. ferrugineus, D. insulanus, D. nicholsonii, D. norrisii, D. occidentalis, D. revolutus, D. rigidulus, D. tophaceus, D. umbrosus, D. vinealis, Triquetrella californica
CD. Cell lumina rounded to elliptic; papillae either numerous and obscuring the cell outlines, or few and sharp, sometimes inconspicuous; axillary hairs pale throughout; apices and stereids various .....Barbula: B. convoluta, B. ehrenbergii, B. eustegia, B. orizabensis not known from CA, B. unguiculata

CE. Immediate leaf marginal cells long and narrow in one or more rows abruptly set off from the shorter cells within .....CF
CE. Leaves elimbate or with an intramarginal limbidium, or with the limbidium of short cells differentiated in cell thickness .....CR

CF. Leaves arranged in two vertical ranks with the leafy plant thus appearing flat; leaf or its base with the lamina split into two flaps that pocket the stem like the base of an Iris leaf .....Fissidens: F. bryoides, F. crispus, F. curvatus, F. minutulus, F. sublimbatus, F. ventricosus
CF. Leaves mostly spirally inserted (if distichous, without the ampliate-clasping aspect of a Fissidens) .....CG

CG. Limbidium intramarginal, several cells thick, with short cells like those of the median lamina to the outside of the elongate cells .....Amblystegium: Limbella fryei not known from CA
CG. Limbidium marginal, often only one cell in thickness .....CH

CH. Plant dendroid with the deltoid-triangular, almost echlorophyllose stem leaves contrasting sharply with the ovate, strongly chlorophyllose branch leaves .....Plagiomnium: Leucolepis acanthoneura
CH. Plant not dendroid .....CI

CI. Plant very small, with sporophyte immersed and capsule almost globose; growing as a winter and spring ephemeral on open, mostly disturbed soil .....Acaulon: Aphanorrhegma serratum not known from CA
CI. Plant small to large but with sporophyte exserted and cylindrical; time and duration of growth various .....CJ

CJ. Margin geminate-dentate .....CK
CJ. Margin entire or with the teeth singly placed .....CL

CK. Leaves with vertical photosynthetic lamellae on costa; lamina frequently spinose-dentate on surface .....Atrichum: all Atrichum
CK. Leaves without photosynthetic lamellae; lamina never with surface teeth except as associated with injury during development .....Mnium: M. ambiguum, M. arizonicum, M. blyttii, M. marginatum, M. spinulosum, M. thomsonii

CL. Median leaf cells primarily quadrate to short rectangular; leaves strongly comose on the short stem; capsule sulcate with operculum dome-shaped .....CM
CL. Median leaf cells primarily elongate-hexagonal; capsules smooth with operculum apiculate .....CO

CM. Capsules inclined, often sulcate; endostome and exostome present .....Funaria: Funaria hygrometrica, F. microstoma, F. muhlenbergii
CM. Capsules erect, smooth; endostome usually reduced; exostome present or absent .....CN

CN. Exostome present, often inconspicuous (if possible, remove operculum from young capsules); exothecial cells oblong; calyptra cucullate, loosely inflated around capsule .....Entosthodon: E. attenuatus, E. bolanderi, E. californicus, E. drummondii, E. fascicularis not known from CA, E. kochii, E. planoconvexus not known from CA, E. rubrisetus, E. tucsoni
CN. Peristome absent; exothecial cells irregular, mostly somewhat hexagonal; calyptra mitrate to campanulate .....Physcomitrium: P. californicum, P. collenchymatum, P. hookeri, P. pygmaeum not known from CA, F. pyriforme

CP. Median laminal cells elongate-hexagonal to rectangular with angular end-walls, mostly more than 4:1; leaves of dorsal portion of leaning stems markedly narrower and shorter than more lateral leaves .....Pohlia: Epipterygium tozeri
CP. Median laminal cells mostly shorter than 4:1; leaves of erect stems without a dorso-lateral heterophylly .....CQ

CQ. Leaves broadly rounded to apiculate at apex, 1.5-4:1, mostly more than 3 mm long; margin multistratose at least in part .....Rhizomnium: R. glabrescens, R. gracile not known from CA, R. magnifolium, R. nudum not known from CA, R. pseudopunctatum, Mnium blyttii
CQ. Leaves with apices mostly acute to acuminate, if rounded to apiculate, with margin unistratose or leaves smaller or more narrow .....Bryum: B. algovicum, B. amblyodon, B. arcticum not known from CA, B. badium, B. bimum, B. caespiticium, B. calobryoides, B. canariense, B. capillare, B. cyclophyllum, B. elegans, B. erythroloma, B. flaccidum, B. knowltonii not known from CA, B. laevifilum, B. lisae, B. meesioides, B. pallens, B. pallescens, B. pseudotriquetrum, B. schleicheri not known from CA, B. torquescens, B. turbinatum, B. uliginosum, B. weigelii, Brachymenium exile not known from CA, Brachymenium systylium not known from CA, Roellia roellii

CR. Leaves two-ranked, keeled at the base, mostly ampliate clasping at the base in the manner of an iris stem .....Fissidens: F. adianthoides, F. aphelotaxifolius, F. asplenioides not known from CA, F. dubius, F. fontanus, F. grandifrons, F. osmundioides not known from CA, F. pauperculus, F. taxifolius, F. taylorii, Bryoxiphium norvegicum not known from CA, Distichium capillaceum, D. inclinatum, Schistostega pennata not known from CA
CR. Leaves inserted in more than 2 ranks, mostly not obviously ranked .....CS

CS. Leaves with adaxial vertical photosynthetic lamellae or filaments over the costa or over both the costa and the lamina .....CT
CS. Leaves without such adaxial lamellae .....CZ

CT. Leaves mostly very coarsely toothed; if with entire leaves those leaves mostly more than 3 mm long .....CU
CT. Leaves entire to serrulate; plant and leaves much smaller .....CX

CU. Photosynthetic lamellae mostly fewer than 12, covering only a small portion the upper leaf lamina; plants strongly crispate when dry ..... Atrichum, Bartramiopsis lescurii not known from CA, Oligotrichum aligerum not known from CA, O. hercynicum not known from CA, O. parallellum not known from CA
CU. Photosynthetic lamellae mostly more than 20, covering the broadened costa on nearly all the upper leaf; plants mostly not very crispate .....CV

CV. Distal cells of photosynthetic lamellae papillose or cuticular papillose .....Pogonatum: Meiotrichum lyallii, Pogonatum dentatum not known from CA, Pogonatum urnigerum not known from CA, Polytrichastrum alpinum
CV. Distal cells of photosynthetic lamellae smooth .....CW

CW. Leaves strongly crispate when dry .....Pogonatum: Pogonatum contortum
CW. Leaves not at all crispate .....Polytrichum: Polytrichum commune, P. formosum, P. juniperinum, P. longisetum, P. piliferum, P. sexangulare, P. strictum

CX. Leaf margins erect from near the base, cucullate at apex; leaves mostly without hyaline awns; cushion of photosynthetic filaments seemingly filling most of leaf concavity .....Aloina: A. aloides not known from CA, A. bifrons, A. rigida, A. roseae not known from CA
CX. Leaf margins recurved or erect but not at all cucullate; leaves almost always with hyaline awns; photosynthetic filaments or lamellae not obscuring the bulk of the leaf lamina .....CY

CY. Leaves deeply concave with margins almost never with recurvature and with 3-5 photosynthetic lamellae narrowly inserted over costa .....Pterygoneurum: P. californicum, P. lamellatum, P. ovatum, P. subsessile
CY. Leaves not concave with margins recurved near base or nearly to apex; photosynthetic filaments present over costa .....Crossidium: C. aberrans, C. crassinerve, C. seriatum, C. squamiferum

CZ. Plants very small and essentially stemless with the closely crowded leaves forming a bulbiform cluster; costa percurrent to excurrent with the distal cells of the lamina and costa forming a somewhat bleached to almost hyaline awn; leaf cross-section with the abaxial walls of the laminal cells markedly more thick-walled than those of the adaxial surface .....Stegonia: S. hyalinotricha, S. latifolia, S. pilifera
CZ. Plants not in all respects as above .....DA

DA. Leaves with hyaline awns .....DB
DA. Leaves without hyaline awns or with the distal half of the leaf bleached and hyaline, thus appearing awned .....DE

DB. Costa filling half or more of leaf base .....Campylopus: C. atrovirens, C. introflexus, C. pilifer not known from CA, C. schmidii
DB. Costa more narrow .....DC

DC. Plant without sporophytes .....Grimmia: G. alpestris, G. americana not known from CA, G. anodon, G. arcuatifolia, G. caespiticia, G. donniana not known from CA, G. elatior not known from CA, G. hartmanii not known from CA, G. incurva not known from CA, G. laevigata, G. leibergii, G. lesherae, G. lisae, G. longirostris, G. mariniana, G. montana, G. moxleyi, G. nevadensis, G. orbicularis, G. ovalis, G. pilifera not known from CA, G. plagiopodia, G. poecilostoma, G. pulvinata, G. reflexidens, G. serrana, G. shastae, G. tergestina, G. torquata, G. trichophylla, G. ungeri, Coscinodon calyptratus, Jaffueliobryum raui, J. wrightii Schistidium cinclidodonteum, S. confertum, S. flaccidum, S. tenerum
DC. Plant with sporophytes .....DD

DD. Capsules immersed and systylious .....Schistidium: S. cinclidodonteum, S. confertum, S. flaccidum, S. tenerum
DD. Capsules mostly exserted, never systylious .....Grimmia: G. alpestris, G. americana not known from CA, G. anodon, G. arcuatifolia, G. caespiticia, G. donniana not known from CA, G. elatior not known from CA, G. hartmanii not known from CA, G. incurva not known from CA, G. laevigata, G. leibergii, G. lesherae, G. lisae, G. longirostris, G. mariniana, G. montana, G. moxleyi, G. nevadensis, G. orbicularis, G. ovalis, G. pilifera not known from CA, G. plagiopodia, G. poecilostoma, G. pulvinata, G. reflexidens, G. serrana, G. shastae, G. tergestina, G. torquata, G. trichophylla, G. ungeri, Coscinodon calyptratus, Jaffueliobryum raui, J. wrightii, Schistidium cinclidodonteum, S. confertum, S. flaccidum, S. tenerum

DE. Leaves more than 10:1, with the costa percurrent to excurrent, often filling the long and narrow apex; plant erect hardly at all branched .....DF
DE. Leaves broader or the costa shorter .....DY

DF. Leaves falcate-secund, plants green to brownish-green, mostly more than 1 cm tall .....DG
DF. Leaves not falcate-secund; or if somewhat falcate, with older plants densely coated with red-brown rhizoid and with leaves having short basal plications .....DK

DG. Plant irregularly but frequently branched, restricted to frequently flooded logs and tree bases along streams .....Dichelyma: Dichelyma falcatum not known from CA, D. uncinatum
DG. Plant unbranched except for innovations; growing in various habitats .....DH

DH. Outer cortical cells of stem strongly inflated and therefore abruptly differentiated from adjacent inner cells; plant mostly with a hint of falcate-secund expression .....Orthodontium: O. gracile, O pellucens
DH. Outer cortical cells not so enlarged and differentiated .....DI

DI. Alar cells inflated, often red-brown; cells of alar region bistratose .....Dicranum: Dicranum fragilifolium not known from CA, D. howellii, D. majus not known from CA, D. pallidisetum not known from CA, D. polysetum not known from CA, D. rhabdocarpum not known from CA, D. scoparium D. spadiceum not known from CA
DI. Alar cells not inflated; cells of alar region unistratose .....DJ

DJ. Costa without stereids in cross-section; plant primarily growing on rocks mainly in alpine regions .....Kiaeria: Arctoa fulvella, A. hyperborea not known from CA, Blindia acuta, Kiaeria blyttii, K. falcata, K. starkei
DJ. Costa with stereids in cross-section; plant primarily growing on soil .....Dicranella: D. crispa, D. heteromalla, D. howei, D. pacifica, D. rufescens, D. subulata, D. varia not known from CA, Cynodontium jenneri, C. tenellum

DK. Costa filling half or more of the leaf base .....DL
DK. Costa more narrow .....DM

DL. Leaves very narrowly linear-setaceous; leaf margin plane; rhizoids abundant and papillose; axillary hairs persistent, with basal cells of those hairs vividly red to red-brown .....Meesia: Leptobryum pyriforme
DL. Leaves mostly lanceolate; leaf margin erect; rhizoids few and smooth; axillary hair fugacious, never with reddened basal cells .....Campylopus: Atractylocarpus flagellaceus, Campylopodiella stenocarpa, Campylopus pyriformis, C. subulatus, Paraleucobryum enerve not known from CA

DM. Leaves crispate when dry .....DN
DM. Leaves not crispate .....DP

DN. Leaf margins entire, or with a few serrulations at immediate apex .....Dicranoweisia: D. cirrata, D. contermina
DN. Leaf margins serrate to dentate even below the immediate apex .....DO

DO. Leaf margins dentate with teeth mostly composed of more than one cell .....Ptychomitrium: P. gardneri, P. serratum not known from CA
DO. Leaf margins serrate, leaves more than 10:1 .....Anacolia: Bartramia halleriana not known from CA, B. pomiformis, Plagiopus oederianus not known from CA

DP. Leaves distichous with base erect and limb reflexed .....Ditrichum: Distichium capillaceum, D. inclinatum
DP. Leaves spirally inserted .....DQ

DQ. Leaves strongly squarrose with the sheathing base abruptly widened relative to the limb .....Dicranella: D. crispa, D. grevilleana not known from CA, D schreberiana
DQ. Leaves not squarrose with limb and base not strongly differentiated .....DR

DR. Leaf tips fragile with most older leaves lacking those apices .....Dicranum: Orthodicranum tauricum
DR. Leaf tips not so fragile .....DS

DS. Leaf margins recurved .....DT
DS. Leaf margins plane to incurved .....DU

DT. Older stems densely coated with papillose rhizoids; leaf once plicate on each side of the extreme base with that plica reflected in the leaf buttress .....Anacolia: A. baueri, A. menziesii
DT. Rhizoids smooth, not densely coating older stems; leaf not plicate or with plica not so reflected in the leaf buttress .....Ditrichum: D. ambiguum, D. pusillum

DU. Outer cortical cells of stem strongly inflated and therefore abruptly differentiated from adjacent inner cells; plant of logs and trees in the Sequoia forests .....Orthodontium: O. gracile, O pellucens
DU. Outer cortical cells mostly not enlarged as an hyaloderm, or if somewhat differentiated; plant growing on soil in open areas .....DV

DV. Capsules sessile and cleistocarpous; plant of disturbed or exposed soil .....Bruchia: Pleuridium acuminatum, P. subulatum
DV. Capsules exserted, often stegocarpous .....DW

DW. Costa excurrent; sporophytes without a well-defined hypophysis .....Ditrichum: D. flexicaule not known from CA, D. heteromallum, D. montanum, D. schimperi
DW. Costa percurrent in a somewhat blunt apex .....DX

DX. Plant very small, growing on rock; hypophysis not obvious .....Seligeria: Brachydontium olympicum not known from CA, B. trichodes not known from CA, Campylostelium saxicola not known from CA, Seligeria campylopoda not known from CA, S. donniana not known from CA, S. recurvata not known from CA, Tetrodontium brownianum not known from CA, T. repandum not known from CA
DX. Plant an ephemeral of open soil; sporophytes with an hypophysis at least as long as the urn .....Bruchia: B. bolanderi, B. flexuosa, Trematodon boasii

DY. Plant erect and essentially unbranched, or if branched, cells isodiametric or elongate with strongly sinuose walls .....DZ
DY. Plant extensively branched, mostly with main stems prostrate; cells mostly elongate .....FH

DZ. Leaves on both the stems and branches squarrose .....EA
DZ. Leaves not squarrose or only the stem leaves squarrose .....EC

EA. Plant aquatic in rapidly flowing streams, leaves and stems dark black .....Scouleria: S. aquatica, S. marginata
EA. Plant of wetland sites but not in rapidly flowing water; plants green .....EB

EB. Leaves very obviously tristichous; leaf base erect-spreading .....Meesia: M. triquetra
EB. Leaves spirally inserted with sheathing base and squarrose limb .....Dicranella: D. palustris

EC. Costa of mature leaf filling 1/3 or more of leaf base .....ED
EC. Costa more narrow .....EE

ED. Leaves blunt at apex; margin recurved; plant of bogs .....Meesia: M. uliginosa
ED. Leaves acute by an excurrent costa that fills the subula; margin erect; plant of soil in open forests and savannahs .....Campylopus: C. subulatus, Dicranodontium denudatum not known from CA

EE. Leaves strongly obovate-spatulate with distal portion of leaf 4-6 times as broad as leaf base; basal margin of leaf with long cilia .....Tayloria: Oedipodium griffithianum not known from CA
EE. Leaf apex more narrow relative to base; basal margin of leaf without cilia .....EF

EF. Capsules erect with a vertically or laterally expanded hypophysis; leaves somewhat broadened near the apex with cells large and thin-walled .....Tayloria: Splachnum ampullaceum not known from CA, S. sphaericum not known from CA, S. vasculosum not known from CA, Tayloria hornschuchii not known from CA, Tayloria lingulata not known from CA, Tayloria serrata not known from CA, Tetraplodon mnioides not known from CA
EF. Capsules and hypophysis otherwise; leaf shape and nature of cells various .....EG

EG. Leaf margin papillose-crenate; leaves somewhat spatulate .....Tortula: Leptophascum leptophyllum
EG. Leaf margin without papillose crenations; leaves mostly broadest below the middle .....EH

EH. Plants with stems almost absent and with the leaves closely arranged in a bud-like cluster; leaves broadest at the middle or below .....EI
EH. Plant with an obvious stem; leaf arrangement and shape various .....EJ

EI. Capsule sessile and cleistocarpous .....Acaulon: A. rufescens, A. triquetrum; Archidium alternifolium, Archidium donnellii not known from CA, Lorentziella imbricata Physcomitrella patens, Physcomitrella readeri, Physcomitrium immersum not known from CA
EI. Capsule erect on a seta about 10 mm long, stegocarpous .....Pohlia: Discelium nudum

EJ. Plants strongly julaceous with leaves deeply concave and closely inserted .....Pohlia: Anomobryum julaceum
EJ. Plants not julaceous .....EK

EK. Median and upper cells of leaf isodiametric or very short rectangular, often transversely elongate .....EL
EK. Median and upper cells of leaf predominantly more than 1.5:1 .....EZ

EL. Basal cells not differentiated from median cells, all cells thick-walled and isodiametric or nearly so, with rounded lumina; most plants with very broad apical leaves forming a cup in which multicellular, disciform gemmae are placed; plant of logs and tree bases, typically with a rust-red cast in older portions .....Tetraphis: T. geniculata not known from CA, T. pellucida
EL. At least the basal or alar cells elongate and differentiated from median cells; plants without gemma cups; plants without a rust-red cast except for occasional dense rhizoidal cushions .....EM

EM. Leaf bases sheathing or rather tightly wrapped around stem with cells of sheath markedly longer than the nearly isodiametric cells of the limb; leaf limbs lanceolate and rather coarsely dentate .....Oncophorus: O. virens, O. wahlenbergii
EM. Leaves not sheathing, only gradually contracted to the very narrowly ovate base .....EN

EN. Leaf margins coarsely serrulate to dentate with at least some of the teeth composed of 1/2 or more of a cell .....Ptychomitrium: P. gardneri, P. serratum not known from CA, P. sinense not known from CA
EN. Leaf margins entire or occasionally serrulate to crenulate near apex .....EO

EO. Leaves very obviously tristichous; cells somewhat elongate, smooth; plant of bogs .....Meesia: M. triquetra
EO. Leaves not tristichous; cells isodiametric and somewhat mammillose; plant of soil and rock outcrops .....EP

EP. Costa with abaxial lamellae several cells high .....Grimmia: G. ramondii
EP. Costa without abaxial lamellae .....EQ

EQ. Leaves bordered by a row of cells abruptly different in size, wall thickness, cell papillosity or pigmentation from the adjacent internal laminal cells .....Scouleria: Crumia latifolia, Scopelophila cataractae
EQ. Leaves not so bordered .....ER

ER. Adaxial cells of costa at and above mid-leaf rectangular with at least some sinuosity on the lateral walls; leaves asymmetric with rows of cells on each side of costa differing in number .....ES
ER. Adaxial cells of costa quadrate or rectangular, not sinuose on lateral walls; leaves symmetric with rows of cells on each side of costa equal in number .....EV

ES. Leaves bistratose throughout .....ET
ES. Leaves mostly unistratose near costa at mid-leaf, often bistratose in several rows near margin .....EU

ET. Leaves narrowly lanceolate above an abruptly broadened base, often falcate-secund .....Grimmia: G. hamulosa G. unicolor not known from CA
ET. Leaves without a differentiated broadened base, never falcate-secund .....EU

EU. Leaves with numerous quadrate cells at the alar region; sporophytes systylious and immersed; plant primarily of seasonally wet to submerged sites, often in sheet drainage; plants typically with a reddish wash .....Schistidium: S. agassizii, S. atrichum, S. cinclidodonteum, S. dupretii, S. maritimum, S. occidentale, S. platyphyllum, S. rivulare
EU. Leaves with few or no quadrate cells in the alar region, instead with these cells mostly rectangular with thickened end-walls; sporophyte immersed or exserted but never systylious; plant primarily of rock faces and boulders, generally dry; coloration typically green to black .....Grimmia: G. anomala, G. hartmanii not known from CA, G. mollis, G. serrana, G. unicolor not known from CA

EV. Leaves elliptic to elliptic-ovate, rather broad to the apex; costa with only an abaxial stereid with that band elliptic in outline .....Tortula: T. californica, T. leucostoma, T. plinthobia, T. protobryoides, T. systylia
EV. Leaves lanceolate, often narrowly so, if broader, then costa without stereid bands .....EW

EW. Leaf margins or all of upper leaf bistratose .....EX
EW. Leaves unistratose throughout .....EY

EX. Costa cross-section without stereid bands; plant forming low red cushions or tufts on siliceous rocks; leaves not crispate when dry; capsules without an operculum, dehiscent by longitudinal lines .....Andreaea: A. blyttii, A. heinemannii A. rothii, A. schofieldiana
EX. Costa with stereid bands; plant primarily on logs and tree bases near the coast; leaves crispate when dry; capsules with an operculum .....Dicranoweisia: D. cirrata

EY. Leaf margins with at least a few minute serrulations near apex; costa with two obvious stereid bands; capsules with seta more than 10 mm long, purple to red, occasionally yellow .....Ceratodon: C. purpureus, C. stenocarpus
EY. Leaf margins strictly entire; stereid bands absent; capsules immersed to emergent on a shorter seta, not strumose but often sulcate and strangulate .....Orthotrichum: O. euryphyllum, O. holzingeri, O. pumilum, Zygodon menziesii

EZ. Leaves falcate-secund with strongly differentiated quadrate and inflated alar cells; median laminal cells with thick and porose walls; distal portion of leaf somewhat undulate .....Dicranum: D. undulatum
EZ. Leaves not or only slightly falcate, without such an alar region; median laminal cells mostly thin, hardly or not at all porose .....FA

FA. Leaves spreading from an erect base; leaf margins entire or with a few serrulations near apex; median leaf cells rectangular with nearly parallel lateral walls .....Meesia: M. longiseta
FA. Leaves erect to spreading but without a differentiation of the base; leaf margins various; median leaf cells mostly rhomboidal to hexagonal but with longitudinal walls not parallel .....FB

FB. Median laminal cells narrowly rectangular with oblique end-walls, mostly less than 10 µm in width; margin crassiserrulate at least near apex; capsule erect, with neck as long as urn .....Bruchia: B. bolanderi
FB. Median laminal cells rectangular to hexagonal or rhomboidal, mostly more than 10 µm in width; margin various, often dentate or serrate; capsule mostly inclined to horizontal or pendulous, never with such a long and well-defined neck .....FC

FC. Leaves with a hint of falcation, blunt at apex with margins recurved and somewhat crenulate; median leaf cells rectangular, less than 8 µm wide, thick-walled with lumen: wall ratio less than 6:1 .....Dicranella: D. hilariana
FC. Leaves mostly symmetrical with apices and margins various; median leaf cells larger, rectangular or hexagonal with thin walls .....FD

FD. Median leaf cells primarily quadrate to short rectangular; leaves strongly comose on the short stem; capsule mostly sulcate with operculum dome-shaped; teeth of leaf margin when present somewhat vesicular-inflated; perigonia borne on small but elongate branches inserted at base of female branches; axillary hairs of markedly enlarged diameter toward distal end .....FE
FD. Median leaf cells primarily elongate-hexagonal; capsules smooth with operculum apiculate; teeth of leaf margin formed of equally thick-walled cells or of cells thickened at the tooth apex (crassi-serrate); perigonial position various but seldom at the end of a branch inserted at base of female branch; axillary hairs typically of equal diameter throughout .....FG

FE. Capsules inclined, often sulcate; endostome and exostome present .....Funaria: F. hygrometrica, F. microstoma, F. muhlenbergii
FE. Capsules erect, smooth; endostome usually reduced; exostome present or absent .....FF

FF. Exostome present, often inconspicuous (if possible, remove operculum from young capsules); exothecial cells oblong; calyptra cucullate, loosely inflated around capsule .....Entosthodon: E. attenuatus, E. bolanderi, E. californicus, E. drummondii, E. fascicularis not known from CA, E. kochii, E. planoconvexus not known from CA, E. rubrisetus, E. tucsoni
FF. Peristome absent; exothecial cells irregular, mostly somewhat hexagonal; calyptra mitrate to campanulate .....Physcomitrium: P. californicum, P. collenchymatum, P. hookeri, P. pygmaeum not known from CA, F. pyriforme

FG. Leaves mostly with cells more than 4:1; costa usually with scattered prorate cells on some of the distal portion of its abaxial surface .....Pohlia: all Pohlia, all Mielichhoferia
FG. Leaves mostly with cells shorter; costa almost never with abaxial prorations .....Bryum: B. alpinum, B. argenteum, B. barnesii, B. bicolor, B. blindii, B. gemmascens, B. gemmiferum, B. gemmilucens, B. gemmiparum, B. klinggraeffii not known from CA, B. lanatum, B. microerythrocarpum, B. miniatum, B. muehlenbeckii, B. pyriferum, B. radiculosum, B. rubens, B. subapiculatum, B. tenuisetum, B. violaceum, Brachymenium spirifolium not known from CA, Plagiobryum zierii not known from CA

FH. Plant plumose to dendroid, with stem erect or upwardly arching .....FI
FH. Plant not dendroid or plumose .....FP

FI. Paraphyllia or paraphyllia-like rhizoids abundant on erect axis and larger branches .....FJ
FI. Paraphyllia absent .....FL

FJ. Leaves complanate and strongly undulate .....Neckera: Metaneckera menziesii
FJ. Leaves neither complanate nor undulate .....FK

FK. Leaves coarsely dentate at least above the middle; plant dendroid, mostly on soil; paraphyllia-like rhizoids with uniseriate cell arrangement present in parallel streaks along stem and branches .....Neckera: Climacium dendroides not known from CA
FK. Leaves entire to serrulate; plant plumose, epiphytic; paraphyllia simple and unlobed or with lobes mostly more than one cell wide .....Antitrichia: Alsia californica

FL. Stem rigidly erect with branches radiating from near its apex; stem leaves small, deltoid-triangular, bleached and scale-like, radically different from the larger ovate to ovate-lanceolate branch leaves .....Plagiomnium: Leucolepis acanthoneura
FL. Stem mostly arching upwards from the substratum; stem leaves not radically differentiated as bleached scales .....FM

FM. Leaves strongly rugose, wet or dry; median cells mostly 6-10:1 .....Hylocomium: Rhytidium rugosum not known from CA
FM. Leaves not rugose; median cells isodiametric to short rectangular .....FN

FN. Median cells isodiametric to short rectangular or rhomboidal; leaves weakly or not at all plicate .....Neckera: Bestia longipes, Bryolawtonia vancouveriensis, Isothecium cardotii, I. cristatum, Thamnobryum neckeroides
FN. Median cells mostly more than 5:1; leaves at least somewhat plicate, at least on the stem .....FO

FO. Stem leaves strongly spreading to squarrose, broadly decurrent, markedly broader and less decurrent than branch leaves .....Kindbergia: K. oregana, K. praelonga
FO. Stem and branch leaves rather similar to one another, auriculate but not decurrent; median cells very strongly pitted .....Homalothecium: Trachybryum megaptilum

FP. Alar cells strongly inflated in a patch about as long as wide; plant of very moist to submerged sites .....FQ
FP. Alar cells not so inflated; habitat various .....FS

FQ. Costa ending at base of acumen or percurrent .....Drepanocladus: D. aduncus, D. capillifolius, D. polycarpos, Campylium polygamum, Cratoneuron filicinum, Palustriella commutata, P. falcata not known from CA, Warnstorfia exannulata, W. fluitans
FQ. Costa not so strong .....FR

FR. Leaf apex rounded to bluntly obtuse .....Calliergon: C. cordifolium not known from CA, C. giganteum not known from CA, Sarmentypnum sarmentosum not known from CA, Straminergon stramineum
FR. Leaf apex acute to acuminate .....Brachythecium: B. nelsonii, B. rivulare

FS. Quadrate alar cells very numerous, concolorous and thick-walled; plant plumose-erect from a prostrate rhizome; paraphyllia present .....Antitrichia: Alsia californica
FS. Quadrate alar cells otherwise; plant mostly prostrate or weakly ascending; paraphyllia various .....FT

FT. Costa extending at least to base of acumen .....FU
FT. Costa mostly ending near middle to 3/4 .....FW

FU. Leaves strongly squarrose with acumen well-demarcated and channeled, and with leaf margin essentially entire .....Campylium: C. chrysophyllum, C. radicale not known from CA
FU. Leaves not squarrose .....FV

FV. Leaves mostly less than 1.0 mm long, minutely serrulate above, sometimes serrate in the peri-alar region .....Amblystegium: Amblystegium varium, Conardia compacta, Hygroamblystegium tenax
FV. Leaves larger, serrulate to serrate but without the special group of teeth in the peri-alar region; rhizoids smooth, not bearing gemmae .....FW

FW. Leaves narrowly ovate-lanceolate to lanceolate, more than 4:1; leaves mostly plicate .....FX
FW. Leaves broader; leaves plicate or not .....FY

FX. Leaves serrulate to serrate at least near the apex; stem never with a hyaloderm; costa of at least some branch leaves ending in an abaxial spine .....Homalothecium: H. aeneum, H. arenarium, H. fulgescens, H. nevadense, H. nuttallii, H. pinnatifidum, Tomentypnum nitens not known from CA
FX. Leaves entire or nearly so; if somewhat serrulate, with a prominent hyaloderm in the stem cross-section; costa never with an abaxial spine .....Drepanocladus: D. sordidus, Hamatocaulis vernicosus, Leptodictyum riparium, Sanionia uncinata

FY. Median cells typically less than 5:1 .....FZ
FY. Median cells typically longer .....GD

FZ. Leaves bluntly rounded to obtuse at the serrulate to coarsely dentate apex .....Neckera: Bestia longipes, Homalia trichomanoides not known from CA, Porotrichum bigelovii
FZ. Leaves acute to acuminate, or apex entire to serrate .....GA

GA. Quadrate or isodiametric alar cells very numerous (more than ten along margin and toward costa); leaves serrate at least in the distal 1/2 .....GB
GA. Quadrate alar cells fewer; leaves entire to serrate .....GD

GB. Costa forked or spurred near base, broad and flat at base with nematogons on its abaxial surface; margins narrowly and evenly recurved from base to near the apex .....Antitrichia: Antitrichia californica, A. gigantea
GB. Costa not forked or forked distally only; margins plane or recurved only near the base .....GC

GC. Plant with leaves with coarsely serrate to fimbriate leaf margins, less than 1 mm long; terminal cell of leaves abruptly several times as long as the adjacent subterminal ones .....Fabronia: F. ciliaris, F. pusilla
GC. Plant markedly larger with leaves serrulate with alternating large and smaller serrulations along the margin; terminal cell of leaves not differentiated .....Isothecium: I. cristatum, I. myosuroides, I. obtusatulum

GD. Plant julaceous .....GE
GD. Plant not julaceous .....GF

GE. Leaves with the adaxial surface of the costa base covered with short and thin-walled cells similar to those of the adjacent basal cells; leaf base typically not at all decurrent .....Scleropodium: Cirriphyllum cirrosum not known from CA, Pseudoscleropodium purum, S. californica, S. cespitans, S. colpophyllum, S. julaceum, S. obtusifolium, S. touretii
GE. Leaves with all surface cells of the basal, adaxial costa surface elongate and similar to such surface cells throughout the costa; leaves decurrent .....Brachythecium: B. collinum, Pseudoscleropodium purum

GF. Leaves serrate from base to apex, mostly with a blunt apex; operculum long rostrate .....GG
GF. Leaves entire to serrulate or serrate only at extreme apex .....GI

GG. Plant plumosely branched with stem leaves markedly broader than branch leaves and with those stem leaves flexed away from stem in almost squarrose manner; distal cells of leaves not markedly shorter than proximal cells of leaves .....Kindbergia: K. oregana, K. praelonga
GG. Plant more irregularly branched, almost isophyllous and without squarrose flexing of stem leaves .....GH

GH. Distal cells of leaves markedly shorter than proximal cells of leaves; distal leaf cells not prorate .....Eurhynchium: E. hians, E. pulchellum, E. striatum, Platyhypnidium riparioides, Steerecleus serrulatus
GH. Distal cells of leaves similar to those of proximal region; many distal cells of leaves dorsally prorate .....Brachythecium: B. bolanderi

GI. Leaves broadly rounded, sometimes mucronate at apex; plants of bogs and fens .....Calliergon: Pseudocalliergon angustifolium, P. trifarium not known from CA, P. turgescens not known from CA
GI. Leaves acute to broadly acute at apex; plants of mesic or hydric habitats but not in bogs or fens .....GJ

GJ. Margins plane, entire or sometimes with a few minute serrulations at apex; costa never ending in a spine; dry leaves loosely appressed to spreading; pseudoparaphyllia foliose but with the outermost bract inserted laterally or proximally .....Amblystegium: A. juratzkanum, A. serpens, A. varium, Leptodictyum humile, L. riparium
GJ. Margins typically at least slightly recurved near base; serrulate, near the apex often coarsely so; costa frequently ending in a spine; dry leaves appressed to loosely spreading; pseudoparaphyllia foliose and so arranged that the outermost bract is inserted on the distal side of the cluster .....Brachythecium: B. albicans, B. asperrimum, B. calcareum, B. erythrorrhizon, B. fendleri, B. frigidum, B. holzingeri, B. hylotapetum, B. leibergii, B. occidentale not known from CA, B. oedipodium, B. oxycladon not known from CA, B. plumosum, B. populeum, B. reflexum, B. rutabulum, B. salebrosum, B. starkei, B. velutinum, B. venustum, Cirriphyllum cirrosum not known from CA


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